Mammals
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Mammals
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Wood
echidna (Dendroglossus scandens)
Order: Monotremes (Monotremata)
Family: Echidnas (Tachyglossidae)
Habitat: tropical rainforests of Australia and New Giunea.
Climatic change in Neocene has strongly affected upon life of various Earth
inhabitants. Some of them had to abandon their habitats, other oness had died
out, and third ones had received great opportunities for evolution and had
turned to prospering species. Humidifying of climate has caused reduction of
desert areas and expansion of tropical rainforests zone. Some of inhabitants
of open spaces have to found convenient places for life in forests, and than
evolved to true wood inhabitants. In Australia which even after intervention
of humans has kept an originality of fauna and flora, amazing abilities to
adaptation were shown by echidna - the inhabitant of savannas and light forests.
It had turned to true wood inhabitant, and besides it had changed to the excellent
tree-climber.
To see such animal on the tree is not so such unusual thing as it seems at
first sight. Among tree-climbers in different continents there were kangaroo,
porcupines and even ground tortoises, not to mention toads and frogs. Therefore
the wood echidna of Neocene epoch, sauntering along tree branches, does not
seem as something unusual on their background.
This is rather large animal: its body length is up to 70 cm, heads with long
proboscis is about 30 cm long. Slowly moving in tree crones wood echidna outwardly
resembles a hybrid of bear and anteater. The wood echidna had kept characteristic
shape of these animals, but had got additional features of adaptation to clambering
habit of life. Its paws which are quickly digging off the firm dry ground,
now equally easily break rotten wood or strong cover of termitary. Claws becoming
more bent cling to bark and branches, permitting the animal to make walks at
30-meter height so easily, as if on firm ground. For more tenacious grip thumbs
on forepaws began to oppose partly to fingers. On hinder legs feet are turned
back, but it at all does not prevent this animal to move. Even on the contrary
to it, using such feet it is easier to climb down headfirst along the tree
trunk. The little toe on hinder legs became a certain analogue of the thumb:
it can be opposed to other toes too and serves for branch gripping. This animal
is very strong: by paws the echidna breaks trunks of trees beginning to rot
and turns out from trunk thick boughs, searching for ants and termites.
Body of the wood echidna is massive and heavy-build. The tail of animal is
very short; it serves practically only for fat accumulation.
The muzzle of this echidna is long, tiny mouth opens on its tip. Main food
of animal includes insects, usually ants and termites, and the main instrument
of their catching is the long tongue covered with sticky saliva. Except for
ants and termites, the echidna eats larvae of beetles searching them in pierced
trunks of dead trees, and also insects living among epiphytic plants. The wood
echidna is able even to plunder trap leaves of insectivorous plants, catching
from them insects floating in digestive liquid secreted by leaves. Sometimes
it simply drinks this digestive juice with insects. One more dainty for the
echidna includes bird eggs. Having found a nest, the echidna drives hatching
bird out from it, accurately pierces an egg-shell by claw tip and licks by
tongue the egg completely. The animal exhausts all eggs in nest though sometimes
it is attacked not only by nest owners, but also by their numerous neighbours
have flied to help. But what can birds make with the prickly robber surpassing
them many times in weight?
Eyes of the wood echidna are very small, and auricles are not present at all.
This animal is short-sighted, and the prickly armor perfectly protects it from
any predator of Australian and New Guinean woods, therefore the wood echidna
can not react (at least, externally) to loud alarm cries of wood inhabitants.
But sense of smell and touch at it are very sharp: the animal easily will feel,
as on branch any bug or spider creeps, not seeing it at all.
Colouring of animal began more contrast, than at its ancestor: needles are
ivory-colored, top of body is light, and stomach is dark, covered with short
thin fur. At the wood echidna there is very dense and thick skin, therefore
stings of ants and termites are not harmful to it. Even army ants killing all
alive on their way, do not harm this phlegmatic animal: unless they will inflict
some stings to sensitive proboscis or between toes. The animal has natural
immunity to their poison, therefore the echidna does not avoid their attacks:
such attack does not harm to it, and in it there is even such benefit – ants
at the same time kill blood-sucking insects in wool of the huge echidna. If
insects (and also larger enemies) appear too importunate, the wood echidna
sprinkles out a jet of musk liquid from special glands. Its sickening smell
easily will drive off the most persevering predator.
The wood echidna is real wood tramp. It has no precisely bordered territory,
and does not protect residence from neighbours. Some animals can peacefully
food on the same tree, and even to ravage one termitary in common, not paying
attention against each other. During such meetings pairing happens more often.
Courtship ritual at the wood echidna is not present, and the breeding season
lasts all the year round.
Not changing traditions of ancestors, wood echidna lays eggs. In its clutch
there are 1 - 2 rather large eggs (with small chicken egg-sized ones), covered
with white skinny shell. During formation of eggs because of hormonal influence
the cross skin plica on the stomach of the female is increasing and turning
to brood pouch. The female lays eggs, lying on back so that the tail appeared
above a head. Then the egg rolls down to the pouch itself from oviduct. The
incubating lasts about three weeks. Cubs hatch underdeveloped, looking more
similar to embryons. They lick milk, squeezing from pouch walls. Usually from
two cubs only one cub survives. It grows quickly, and at the age of two months
(when it leaves mother’s pouch), the cub weighs already about half-kilogrann.
It is covered with grey naked skin, but at it the short proboscis characteristic
for these animals is visible. Claws of it still soft and short but after the
cub has left the pouch, it literally starts to increase from different directions:
practically simultaneously at it needles, wool and claws grow. While the cub
is still small, its unique protection is musk liquid with which it excretes
plentifully at a fright. The female regularly visits it (approximately twice
per day) and feeds with milk. For this purpose it lies down on back in forked
crown of branches, and the cub licks off the milk flowing down from pouch to
stomach. But gradually the cub also starts to catch insects. At the age of
three months the cub already starts to wander in wood canopy with mother, studiing
to swarm up lianas and to search for insect nests. At the age of eight months
it reaches length about 70 cm and already is completely independent. At the
age of 20 months the young female already can lay the first egg and this surprising
animal can live up to the age of 60 - 70 years.
Stegoechidna (Stegotachyglossus armatus)
Order: Monotremes (Monotremata)
Family: Echidnas (Tachyglossidae)
Habitat: forests and scrub of Northern and Eastern Meganesia.
During the Cenozoic era the branch of monotreme mammals gradually declined.
Their variety gradually became lesser, and the habitat area narrowed steadily.
In human epoch the area of monotremes included only Australia and New Guinea.
But in these districts monotremes were competitive enough, and strongly kept
their place in ecosystems. Before the occurrence of people in Australia the
sheep-sized giant echidna existed in this continent. But human activity had
quickly resulted in extinction of these animals.
Having survived in human epoch, monotremes had proved their viability. In Neocene
representatives of this group of animals had formed a number of new species.
The apopheosis of evolution of this order of mammals is the largest species
of ever living monotremes, the inhabitant of rainforests at the north and east
of Meganesia. This animal is named stegoechidna.
The appearance of stegoechidna causes memoirs on epoch of dinosaurs. This is
a large four-footed animal, growing to about one and a half meters high at
a shoulder and weighting up to 200 kgs. It has a massive constitution and thick
short tail. Back and tail of this animal bristle up with corneous prickles
and scales, providing reliable protection against an attack of predator. Along
the back of animal pointed and long corneous spikes of reddish color grow,
gradually transforming to reddish plates on tail and hips. The skin of this
animal about 2 sm thick is covered with short and rough black wool. The top
layer of skin cornificates, forming additional protection.
Head of stegoechidna is narrow and long, with slightly curved “Roman” snout
and small mouth. The bottom jaw is mostly long and thin, and only in its basis
bones extend. In these places on bottom jaw thick corneous plates, with which
help the animal crushes forage, develop. At this animal there are small eyes,
and also external auricles are not present. Sight and hearing at stegoechidna
are weak, but sense of smell is very keen.
This animal lives in underbrush and moves, supporting on external part of hand
and having turned up huge forepaw claws. With their help animal digs ground
in searches of forage. As against to its smaller relatives, this animal is
omnivorous, and the significant part of ration of stegoechidna is made of vegetative
food – tubers and roots of plants. Animal digs them out by monstrous claws
of forepaws, takes by small mouth and pounds between ridge corneous plates
growing on jaws and tongue basement. Also it willingly eats earthworms, grubs
and fruits fallen from trees.
Stegoechidna has very small brain, and its behaviour differs in primitiveness.
It leads a solitary way of life, and shows interest to relatives only during
the breeding time.
These animals breed at presence of enough of food, usually in late spring or
in the beginning of summer. The female lays three large eggs with dense shell
into the brooding pouch forming at this time. The incubation lasts about two
months. Newborn cubs eat milky secretions of skin within three months, and
leave a pouch rather well developed. Having left the pouch, they follow the
mother and eat larvae of insects and other invertebrates, yet become advanced
enough to digest vegetative food. Young animals abandon mother at the age of
about 5 months. They reach sexual maturity at the 6-th year of life.
Life expectancy of stegoechidna reaches 60 – 70 years. The adult animal does
not have enemies – only few predators can bite through its thick skin. However,
the young animals have recently left the pouch are protected much more poorly,
than adult individuals – corneous scales and spikes at them develop within
approximately one month after they leave a parental pouch. At this age the
greatest number of animals of this species perishes.
This species of animals was discovered by Tim Morris, Adelaide, Australia.
Starbill (Astrorynchus condyluroides)
Order: Monotremes (Monotremata)
Family: Duck-bills (Ornithorhynchidae)
Habitat: lower reaches of rivers of southeast of Meganesia, Flinders Island
in Eyre Gulf.
Climatic changes in Neocene favour to evolution of thermophilic and moistureloving
species. The most part of territory of Australia in Holocene was covered by
droughty deserts. In Neocene this continent had moved to the north and had
joined to New Guinea forming unite continent Meganesia. Rivers supplying with
water plains of the central part of Meganesia spring from the north and east
of the continent. The Eastern Meganesia prospers due to large river system
occupying the territory of former Murray River. It runs into extensive shallow-water
Eyre Gulf filled with warmed up brackish water. Expansion of habitats stimulated
the evolution of water inhabitants.
The genealogical branch of monotremes declined till the Cenozoic Era had received
new opportunities in Neocene. The occurrence of productive ecosystems has allowed
to evolve to the various specialized species of these primitive mammal. The
large piscivorous duck-bill mergotherium
(Mergotherium piscivorus) inhabits
rivers, and in mountain streams its tiny aggressive relative ancistrotherium
(Ancistrotherium radulus) lives. In lower reaches of rivers running into Eyre
Gulf, and also at small Flinders Island, one more descendant of duck-bill,
the starbill, lives.
This species of duck-bills resembles the American star-nosed mole in many respects
– it leads similar way of life, and even has few common features in appearance.
This is small (rat-sized) animal with powerful forepaws using for digging.
Between toes membranes are advanced; the animal is nice swimmer. At this animal
the swimming membrane on forepaws is reduced, and they are more adapted to
digging, than forepaws of duck-bill. During the swimming starbill presses them
to the body. Besides the starbill is able to climb on roots in mangrove thickets,
and often catches ground insects and snails. The tail at starbill is weak and
short – this animal swims with the help of strong hind paws.
The most original part of appearance of starbill is its beak, the family feature
of duck-bills. Beak at this species is toothless, short and wide. But on it
the set of sensitive leathery outgrowths grows; that’s why the beak of starbill
resembles a little the snout of star-nosed mole (hence the name: Condylura
– the American star-nosed mole).
This animal is protected from predators by poisonous spurs (it is one more
feature inherited from duck-bill) colored bright orange. At males spurs are
longer, than at females. Fur of animal is black or dark brown with contrast
white marks on head and shoulders. The predator attacking starbill will remember
its colouring, having received a painful prick of poisonous spurs.
Starbill lives in damp ground of riverbanks and in marshlands. By this feature
it also resembles its prototype from the insectivores order. It eats various
ground and water invertebrates – earthworms, leeches, snails and larvae of
insects. The animal searches for them with the help of sensitive outgrowths
on edges of beak. These outgrowths are rich in chemo- and electroreceptors,
and permits to define presence of live creatures even in darkness. Under water
the starbill closes eyes, and is guided exclusively with the help of touch
sence and receptors of beak.
Starbill is a solitary animal. Each individual digs complex system of tunnels
in riverbank. The nest for posterity rearing represents the long hole terminating
in the nesting chamber. This chamber is located above maximum water level,
and has an emergency exit.
In lower reaches of rivers starbills of different genders lodge in different
places. Males prefer to live in territories completely filling with water during
the inflow. Here the amount of forage is more, and for spending the night animal
often uses any casual shelter. Females prefer to settle closer to riverbanks
where their long-term holes will not be filled with inflow.
At Flinders Island the separate population of this species differing in smaller
sizes (animals weigh approximately 20% smaller, than their relatives from the
“motherland”) lives. Conditions of life at this island are more severe, than
at the continent – permanent sources of fresh water are not present there.
Therefore starbills of Flinders Island receive necessary moisture exclusively
from food. Also at these animals there is stronger poison secreting in smaller
amount.
This animal was discovered by Simon, the forum member.
Ipochereu,
Amazon marsupial marten (Martenodelphis ipochereu)
Order: Marsupials (Marsupialia)
Family: American marten opossums (Martenodelphidae)
Habitat: tropical woods of Amason region, mountain of an equatorial zone.
Marsupials and placentary mammals are not two levels of development, and two
equivalent groups of mammals. Marsupials do not have attributes because of which
they can be counted “lower” ones. Presence of brooding pouch and way of posterity
bearing has imposed some restrictions to adaptive opportunities of marsupials:
among them forms constantly or the most part of time living in water basically
can not appear. But in ground habitats marsupial mammals rather successfully
can compete with placentary ones. Isolation on southern continents obviously
has not was of use for them: developing “using own sources” and being out of
competition, marsupials lost in ingenuity level to the placentary contemporaries
from other continents. After to South America and Australia placentary mammals
have got, the significant part of marsupials had lost struggle for existence
and had disappeared from Earth’s face. But extinction of plenty of mammal species
in late Holocene and early Neocene had balanced chances of marsupials and placentals
in the further existence. In South America some new kinds of marsupials successfully
competing with placental ones had appeared. Among them representatives of American
marten opossums family, predators of woods and mountains,
are especially remarkable. They had descend from American opossums (Didelphidae),
but evolution has perfected their predating abilities, having made these animals
one of the most successful group of local predators.
In forest canopy of the Amason region, stretched to many thousand square kilometers,
one of representatives of this family, ipochereu, or Amazon marsupial marten
lives (Ipochereu is a name of marten, the hero of fairy tales of one Indian tribe
of Amason region). By constitution ipochereu resembles the true marten, but it
differs from them in larger size: length of body is about 70 cm (tail is about
1 meter long), height at a shoulderis about 30 cm. The long body and short paws
with tenacious sharp claws are adaptation features for life in rich branches.
Amazon marsupial marten differs in big flexibility and mobility: this predator
easily can run across the rivers along thin liana at 30-meter height. Due to
tenacious claws this animal moves on tree trunks easily as if squirrel. The tail
of this marsupial marten is not prehensile, but during jumps from tree to tree
it serves as the balance weight.
Colouring of wool of Amazon marsupial marten is spotty - on grayish-brown background
there are longitudinal lines of small white spots merging on back to faltering
strips. Stomach is white.
Obverse part of skull is short, but jaws are wide and strong. Brain cavity of
skull is long and low. This predator kills catch (large forest rodents, birds
and reptiles) by strong bite to backbone or in basis of skull. Eye-sockets are
wide, sight is substantially binocular due to what the animal can precisely estimate
distance before jump. Ears are short and rounded. On muzzle long whiskers grow.
As the majority of large predators, Amazon marsupial marten is solitary animal.
Each animal occupies extensive territory - about 10 square kilometers. Borders
of territory are marked by urine and areas of toren bark (as bears in Holocene
acted), and carefully protected. Only males during breeding season come to the
territory of females for pairing.
Amazon marsupial marten has two peaks of activity: in the morning (from dawn
up to hottest time of day) and in the evening up to sunset. In the afternoon
in heat the animal has a rest in shadow or hides in tree-trunk hollow. Sometimes
Amazon marsupial marten goes down to the ground and sleeps in shadow under tree
roots. On the ground this predator finds food – carrion, eggs of turtles and
other reptiles. Sometimes ipochereu catches fish and crabs in shallow streams
(especially frequently it makes it in mountain areas).
At ipochereu the expressed seasonal prevalence in breeding is not present, but
in mountains or in the south of area cubs are born in early spring, and even
at the end of winter. The pouch at this species of animals represents the horseshoe-like
plica of skin inverted by the ends to tail. Up to the moment of posterity birth
it is stretched, providing preservation of posterity and when cubs leave it,
pouch is compressed again. In pack there are up to 5 cubs born underdeveloped
as at all marsupials. They stay in pouch within three months, eating milk and
developing. Then the female some time carries them on back (the relationship
with opossums has an effect), but at this species similar feature gradually passes
to past. In mountains some females may at once leave rather developed cubs in
shelter (similarly to Australian predatory marsupial): cubs sitting on back simply
prevent to hunt. Besides sometimes during catch chasing the cub simply can fall
down from mother’s back and injure itself up to death. Because of it before independence
only 1 - 2 cubs from pack survive. At the age of 5 months they start to hunt
with mother, and completely pass to meat feeding. At this time the female is
ready to rear posterity once again. At the age of seven month young animals already
are completely independent, and the female already bears the next pack in brooding
pouch.
Marsupial
sharp-toother (Carnomacropus raptor)
Order: Marsupials (Marsupialia)
Family: Predatory marsupials (Dasyuridae)
Habitat: savannas and light forests of Australia.
Looking at this carnivorous creature, especially during its hunting, it is
possible to think involuntarily, that dinosaurs have returned to Earth.
But actually it is not so, though this animal by habits and constitution is
very similar to reptiles missed for a long time from the face of Earth.
The marsupial sharp-toother is very original animal, and in fauna of Earth
of human epoch there were no analogues to it. It is animal about meter height
at a shoulder and weighting up to 50 kgs, the huge descendant of small marsupial
jerboas (Antechinomys). It has inherited and has developed characteristic carnivorous
habits of ancestors, and increasing the size this animal has had an opportunity
to attack not only small animals, but also to overpower even sheep- or antelope-sized
animals. Marsupial sharp-toothers became original "cheetahs" of the
Australian savanna attacking running herbivores. They are solitaries but if
they have mastered concerted group hunting, it would not be more terrible animal
in Australia than this creature.
The constitution of the marsupial sharp-toother is adapted to fast chasing
of catch: back legs of animal are advanced as well as at kangaroo, and armed
with moderately long non-plungering claws (three claws on each foot). At short
distances the animal can accelerate momentum up to 80 kms per hour, and at
long distance it keeps speed about 40 kms per hour during tens minutes. Forepaws
of animal are rather big, there are advanced long fingers with the sharp claws
on them, permitting to grip and kill small catch. Tail of animal is long (the
common body length is 2 meters, half of which tail amounts), well permits to
turn and to keep balance at sharp turns.
The head of the marsupial sharp-toother is armed with powerful jaws and well
advanced canines: tips of canines are visible from the closed mouth. Incisors
are very sharp, molars are powerful and wide: at lack of alive catch the animal
eats carrion, even dried up at the sun. Frequently marsupial sharp-toothers
eat up the rests of catch of other large predators. The chewing muscles are
well advanced: the animal kills large animals by bite in neck, or inflicts
by teeth extensive wounds to catch and drives it up to exhaustion. Ears of
animal are wide, in them there is advansed the circuit of blood vessels: in
heat weather they serve for heat emission. Nasal cavity is high, nostrils form
a kind of small proboscis – it simultaneously increases olfactory surface and
clears inhaled air of dust: at internal edge of nostrils some numbers of the
hair playing a part of dust filter grow.
Eyes of animal are directed forward, sight is binocular.
Colouring of body of the marsupial sharp-toother is basically sandy-yellow
with faltering pale grey strips on shoulders and head, stomach is lighter,
on tail tip there is a hairy brush of long black hair.
Marsupial sharp-toothers are solitary animals meeting each other only for the
period of pairing. Also the grown up cubs some time keep near to mother, training
in hunting technique. Males at this species are larger and more aggressive
than females. Courtship relations of animals remind fight more: the male forces
the female to pairing by every possible way, bites its skin on back and forepaws.
At adult female’s ears are frequently torn up and bleed: these are traces of
similar “grooming” of males. Pairing repeats some times within two - three
days, then the male abandons the female and does not accept any participation
in posterity rearing. Pregnancy lasts not for long as at all marsupials: about
one week; newborn cubs are underdeveloped, more similar to embryons. Usually
delivery begins at night, when there is no danger of drying of the cub on the
sun. The female lies down in the dug out small hole so that the aperture of
maternal passages is little bit higher than an entrance to the marsupium (which
opens back), and gives rise up to 5 - 8 cubs. But because in pouch there are
only 4 nipples, only those survive who reached to the pouch the first. The
aperture of pouch is surrounded with elastic ring muscle which pulls it during
movement of animal; therefore cubs do not drop out of pouch during sharp throws
at catch chasing. In pouch cubs stay about 3 months, thus from four only two,
and sometimes only one can survive. But its chances of the further survival
are rather great: the female furiously protects cubs, attacking even on the
largest herbivores if it will seem that cubs are threatened with danger. At
the age of four months the cub freely follows mother and can catch small vertebrates,
and one-and-half year aged cub participates in hunting on equal with mother
(as a rule, at this time it already bears a new litter in pouch). Young animals
become sexual mature rather early: one-year-old females continue to grow, but
already can rear posterity.
Hunting small animals, for example, rabbits, the marsupial sharp-toother tries
to creep to them as closely as it is possible under covering of grass and bushes,
and then overtakes catch in short throw. Having overtaken with it, it inflicts
lateral impact by claws of forepaws, trying to knock down and to wound catch,
and then turns around and inflicts to chase a fatal bite. Hnting larger animals,
for example, camelopes (antelope-like camels), the marsupial sharp-toother
separates the planned chase from herd, and then drives it some hours along
the plain, not developing maximal speed. Usually during continuous chasing
the prey weakens so, that the predator succeeds to overtake it by one conclusive
throw and to seize its throat.
Usually marsupial sharp-toother is active since early morning till midday and
in the evening. The animal spends hot daytime in shadow of trees or in rich
bushes. If it is a lot of catch, the animal can any time stay in feeding area.
Then in bushes or river breakage it digs to itself wide hole where it has a
rest in the afternoon and sleeps at night. At lack of catch the animal becomes
the nomade and lives in temporary shelters.
Marsupial
shrew (Nanantechinus aquadromeus)
Order: Marsupials (Marsupialia)
Family: Dasyurids (Dasyuridae)
Habitat: Northern and Eastern Meganesia, river walleys, lakes and bogs.
Till the Cenozoic, before the appearing of man and his satellites from among
placentary mammals, marsupial mammals of Australia had developed practically
all accessible ecological niches, having formed a plenty of various species.
But one environment had remained completely inaccessible for them – water habitats
were those ones. Among Australian marsupials there are no analogues of otters,
seals or beavers. The brooding pouch, the refuge for young growth for long weeks,
had made development of water inhabitancy impossible for marsupials. Therefore
in Cenozoic in Australia the vacant place of large water vertebrates was shared
by birds, turtles, platypuses and rodents.
In Neocene marsupials had made one original attempt to develop water habitats.
Floating plants covering with leaves and stalks a surface of reservoirs, became
the house for one such creature. At the end of day and at the early morning on
the surface of carpet of floating plants it is possible to notice this creature,
managed to find the compromise between presence of brooding pouch and semi-aquatic
habit of life. It is tiny marsupial shrew - one of tiniest mammals of Neocene.
Length of body of this creature may be about 4 cm; tail is about 2 cm long. The
body constitution of marsupial shrew is very graceful; these small mammals are
similar to shrews with long legs. Back legs of this animal are little bit longer
than front ones. Head is rather large, with wide rounded ears. The muzzle is
long, extended to mobile proboscis, on each side of which thin whiskers stick
up. The marsupial shrew is the descendant of one of small species of marsupial
mice (Antechinus).
As this animal lives in areas with a warm climate, its wool is short and rigid.
It is colored light yellow, almost white color – it protects small mammal from
overheat. The body temperature at the marsupial shrew is changeable: at night
the animal is hidden among plants and runs into original catalepsy during which
the body temperature lowers down to ambient temperature. At this time the marsupial
shrew is rather languid, therefore it tries to find a reliable place where it
will be not found by predator. Sometimes for spending the night the animal even
gets into buds of water lilies which are closed for night, but do not immerse
into the water. In hottest time of day marsupial shrew is also hidden in shadow
to avoid overheat, but thus its temperature does not fall.
Fingers and toes of animal are bordered by “brushes” of hairs, and marsupial
shrew greases them with fat secretions of anal glands. Due to it hairs on paws
become not moistened, that permits this tiny creature to live on the surface
of reservoirs. It quickly runs on floating water plants, and even can run small
distances on the surface of water, quickly splashing on it by paws like the basilisk
lizard (Basiliscus) known in Holocene epoch. Basically, young animals and adult
males which are smaller, rather than females approximately to 30 – 40% are able
to do it. Males and females without cubs can even dive for some seconds. The
female, especially one with cubs, does not risk to run on water and to dive.
It keeps on floating plants where it is easier to find forage and to hide from
enemies.
At this creature there are very sharp teeth: marsupial shrew is a gluttonous
predator, and devours for day twice more food, than it weighs itself. It eats
insects and their larvae, and also small shrimps. The animal catches them, immersing
head and forward part of body under water. Due to small size marsupial shrew
catches larvae of mosquitoes in leaf axils of floating plants – this animal alongside
with fishes and predatory insects controls the amount of mosquitoes in damp districts.
Also it can catch fish fry hiding among plants, and even attacks small crabs.
Marsupial shrew lives in conditions where seasonal prevalence of climate rather
feebly marked; therefore it breeds the year round. The brooding pouch at it is
reduced up to two longitudinal plicas in groin area, bordering nipples from sides.
In litter it may be only 3 – 5 cubs developing very quickly. At monthly age they
already creep on back of the female, and in two months abandon mother. One –
two weeks later the female is ready to reproduction again. Sexual maturity at
young animals becomes at the age of three months. Life expectancy of the marsupial
shrew is more, rather than at placentary mammals of similar size with constant
body temperature: it may amount about two years. It is the result of night catalepsy
which is slowing down the “combustion” of its organism.
The idea about the existence of this animal is stated by
Tim Morris,
Adelaide, Australia, at the basis of idea of Dougal Dixon.
Marsupial crab-eater (Cancrivenator dexter)
Order: Marsupials (Marsupialia)
Family: Predatory marsupials (Dasyuridae)
Habitat: zone of mangrove forests separating Arafura Lake from ocean.
Mangrove forests at the northwest of Meganesia became the house for numerous
and various species of animals. There are many species of various mammals among
inhabitants of these forests. They can be vegetarians or predators, but one
feature unites all of them – they perfectly climb on trees. Some of them eat
only the food given by trees, and others feed in swamps, using trees only as
a place for life. The strange descendant of marsupial mice belongs to such
species – it is mobile and dexterous creature named marsupial crab-eater. The
shape the marsupial crab-eater resembles tarsiers (Tarsius) of Holocene epoch,
but it strongly differs from them by habit of life.
Marsupial crab-eater is medium-sized mammal. It weighs about 400 grams and
it seems rather large because of lengthened paws and long flexible tail. Rear
legs of this animal are much longer than front ones, and long tail serves as
the balance weight at jumps. Besides the bottom side of tail is naked, and
tail is also prehensile. The convergence of marsupial crab-eater and many climbing
mammals of New World also had such adaptations ise shown in it. Fingers and
toes of marsupial crab-eater are prehensile and mobile. The animal is able
to jump and to swarm up trees. Moving down from mangrove trees to the swamp,
marsupial crab-eater does not jump off on the ground – it may easily stick
in silt and die. It dexterously skips on roots of mangrove trees only on hind
legs, and climbs on branches in tree crone with the help of all four paws and
tail. Marsupial crab-eater can make long jumps from tree to tree, especially
being pursued by predators.
The wool of this animal has reddish-brown colouring with yellow spots forming
longitudinal lines on back. Throat, breast, stomach, forepaws and the bottom
part of hinder legs are white.
Marsupial crab-eater has large eyes directed forward and providing almost completely
binocular sight. Also it has wide rounded ears and long mobile proboscis-like
nose. Marsupial crab-eater is an exclusive predator with sharp incisors and
multituberculate molars. It eats small animals, searching them on roots of
mangrove trees during the outflow time. The basic prey of this animal includes
crabs; animal prefers to catch young crabs and crabs of small species. It catches
crabs by very original way: in order to seize the crab, the animal overhangs
headfirst and keeps by hinder legs and tail for roots of mangrove trees. By
free forepaws marsupial crab-eater seizes crab for the bases of claws, and
thus does not give it to protect itself, and then it puts to prey fast bite
by powerful incisors. This animal eats prey, sitting vertically on tree branch
and holding it in forepaws. Marsupial crab-eaters drink rain water, licking
it from leaves, or cautiously drinking up the top layer of fresh water from
the surface of mangrove swamp channels.
Marsupial crab-eater is a solitary animal meeting congeners only in courtship
season. These animals declare rights to territory, uttering loud shrill cries.
If two animals meet each other at border of territory, they display force to
each other, having seized branch by all paws and shaking it. After rain animals
mark borders of the territory with urine.
In the most remote place of the territory each animal builds on tree wum sphere-shaped
nest among live branches, bending and plaiting branches with fibres and strong
leaves of plants like pandanus (screw palm). Branches continue growing, and
the nest becomes strong and imperceptible. In this nest animal spends night,
and female leaves the grown up cubs.
The courtship season at marsupial crab-eater proceeds during all year round.
Males of this species search for females ready to pairing by smell. Courtship
at marsupial crab-eater is rough enough and hard: male interferes to the territory
of the female, pursues her and forces to stop for pairing, biting her tail
and hips. At this time he utters the special sounds similar to grumble.
After very short pregnancy at the female up to five underdeveloped cubs are
born. But in pouch opening forward like at kangaroo, there are only two nipples;
therefore only two cubs survive. They develop in pouch for a long time (till
about four months). The grown up cubs still some time sit on back of the female,
keeping for its wool and having clasped its tail. The female leaves grown up
and already heavy cubs in nest and comes back to them for feeding 4 – 5 times
a day. At this time cubs gradually start to study climbing and jumping on trees.
When they become independent enough and start to accompany with the female
during hunting, she has already time to couple and soon gives rise to new cubs.
Marsupial crab-eater becomes sexually mature at the age of one year. Life expectancy
of these animals makes 10 – 12 years.
Filicide embryotherium (Embryotherium infanticida)
Order: Marsupials (Marsupialia)
Family: Predatory marsupials (Dasyuridae)
Habitat: Meganesia, woodlands and forests of various types.
People-caused introducing of placental mammals to Australia had strongly reflected
in evolution of local marsupials. Many carnivorous species of marsupials had
been exterminated by people or had died out, when people had exterminated large
herbivorous marsupials. And small marsupials frequently did not maintain a
competition to placental predators. But one group of Australian predatory marsupials
had got great advantage from occurrence of placentals in Australia – these
ones were small predators, marsupial mice. When epizootias had reduced the
number of placental predators occupied Australia, marsupial predators had not
missed their chance and had returned in new quality – as clever, furious and
easily adapting species. Gradually they had taken a place in new ecosystems
of Australia and further in Meganesia.
The descendant of one Holocene species of marsupial mice, tiny (the total body
length is no more 10 cm, not including short tail) species of predatory marsupial
mammals, had developed the unique habit of life combining predating with original
kind of parasitism. It lives in places where many species of large marsupials
live, and spends a part of life, parasitizing in their brooding pouch. This
animal penetrates into pouches of large marsupial mammals and eats still tiny
newborn cub bearing by them. Then adult animal is attached instead of it to
the nipple and feeds on milk secreting by the female involuntarily becoming
the host of parasite. This animal is named filicide embryotherium.
At embryotherium many infantile features are kept in appearance. The skin of
this mammal is covered by thin soft wool and is completely naked in some places.
In appearance of animal the sharp sexual dimorphism is expressed. Wool of females
is thin and sparse; its color is yellowish-white. Males have richer greyish-yellow
wool with cross brownish strips.
Embryotherium, however, spends a significant part of life out of host animal
pouch. It is connected mainly with search of sexual partner, settling of young
animals and search of host animal by them. Embryotherium has well advanced
sensitive eyes and large ears – not living in host pouch animal leads nocturnal
habit of life and prefers to hide in various shelters in day time. Its sense
of smell is very keen – animal easily distinguishes by smell the female bearing
the newborn cub on the background of smells of other part of herd. Paws of
embryotherium are strong and supplied with well advanced claws. This animal
is rather mobile: embryotherium can swarm up trees, run and jump fast. Its
mobility is connected to necessity of search of host animal.
By the appearance embryotherium resembles a mouse. It has rather large head
with the short muzzle; muzzle, lateral parts of head and throat are almost
hairless. Teeth of animal are short and sharp; only incisors and canines develop.
The reason of it is a fact that till the significant part of life animal eats
soft food and has no necessity to chew it. Milk-teeth of embryotherium are
advanced better, than second ones – it is connected to necessity to lead an
active habit of life till the first weeks of independent life, up to the meeting
of host animal. Some more interesting adaptations to existence in pouch of
host animal are the features of excretory system. Embryotherium produces very
dry dung and concentrated urine. Leaving the nipple of host animal, embryotherium
puts out back from its pouch, sharply throws outside dung and urine, and then
hides again.
Embryotherium is able to hunt and to eat small animals – worms, slugs and soft-bodied
insects like cockroaches and young crickets. Usually such diet is characteristic
for young individuals. Males spend in pouches of host animals rather small
part of life – they grow up, eating milk of the host animal, and at the approach
of sexual maturity leave the safe refuge. Adult males search for females by
smell and pair with them actively. Sperm keeps viability in sexual ducts of
the female till the long time, and it can bring up to three litters after one
pairing. It is an adaptation to secretive parasitic habit of life. After pairing
the female at which cubs develop, searches for the host animal. It settles
in its pouch, kills and eats its cub, sticks to nipple of the host animal,
and raises its own posterity in safety and food abundance. In litter of the
embryotherium it may be up to 6 – 7 cubs. The pouch of embryotherium looks
like two thin and elastic longitudinal plicas surrounding nipples from two
sides. Cubs eat parent milk for very long time – it is their only food till
about 4 months, and during the entire life they keep ability to digest milk
proteins. Young embryotheriums grown up sizable enough, leave host animal’s
pouch and lead independent life within several months. Females reach sexual
maturity rather early – at the second month of independent life. After pairing
embryos in their bodies run to a diapause and do not develop while the female
will reach normal physical standard. Free-living males seldom survive over
two years, but separate individuals succeeded to find the host animal after
rough courtship season, quickly restore forces and can live up to four-year-old
age.
The idea about existence of this species was proposed by Simon and Mutant, forum members.
Marsupial
digger (Pseudotalpa geobia)
Order: Marsupials (Marsupialia)
Family: False marsupial moles (Pseudonotoryctidae)
Habitat: bush savannas of Australia.
Marsupials in Australia have shown to the full an opportunity of evolution
of separate limited group of animals. Among them in different time ecological
analogues of placentary mammals, frequently belonging to different orders,
had appeared. Even the marsupial mole lived in Holocene at the territory of
Australian deserts – the kind of blind digging marsupials.
In Neocene among marsupials the second time the species having underground
habit of life had appeared. It can seem strangely enough, it had evolved due
to activity of human for a long time disappeared on Earth. In due time people
had introduced rabbits to Australia, and they have successfully gone through
cataclysms on border of Holocene and Neocene due to the abilities to adaptation
and high fertility. In rabbit holes spreading under ground to many tens of
meters and forming ramified systems various species of animals settled. Among
them any species of marsupial mice (Antechinus), subsequently adapted to completely
underground way of life was. This small predator had to found in rabbit holes
practically everything necessary for its life: relative safety and food – numerous
insects hiding in holes from day time heat. In holes it was always possible
to find larger catch – dead rabbit cubs. Evolution had picked up and had improved
this idea and as a result savannas of Neocaenic Australia were occupied by
one more species of digging marsupials – marsupial digger.
This animal is medium-sized one: length of body is about 20 cm, and height
at a shoulder is only about 6 – 8 cm. Body of this small mammal is cylinder-shaped
and muscled; it is covered with short velvety brown-colored fur. Paws of marsupial
digger are short, on strong forepaws there are three long strong claws of yellowish
color (on II, III and IV fingers) with which help the animal carves its burrows.
Hinder legs are very short, with narrow feet and four fingers (there is no
thumb at the foot). Tail of this animal is short and very thick; it serves
for accumulation of fat stock. The layer of fat also is laid on waist of animal.
At the marsupial digger there are short wide head and very strong skull. Bones
of skull are thick and also have grown together with each other to solid vault.
Eyes and auricles are absolutely not present at it, and nose is covered with
corneous scute, as at marsupial mole (Notoryctes). Nasal cavity of animal is
strongly expanded, and its internal surface is covered with set of plicas –
animal has keen sense of smell. Looking at teeth of marsupial digger everybody
will understand that it is furious predator in its weight category: all teeth
in its mouth are peaked and cutting; and canines are rather long, even jutting
out from mouth.
The marsupial digger lives exclusively under ground and never leaves on surface.
If the marsupial mole of Holocene epoch preferred to live in drift sands, the
marsupial digger builds constant system of constant holes in dense ground,
in which it waits for catch – every possible small animals. These small mammals
are very strong, and are able to kill animal larger than itself. Metabolism
at marsupial digger is rather intensive: for day it eats quantity of food only
twice less, than it weighs itself. Having found a plenty of food, the animal
eats it all and quickly accumulates fat in back part of body and tail. The
full digger becomes inactive to any time. It creeps to the hole, closes an
entrance by cover of friable ground and vegetative rests, and sleeps. After
the food will be digested, it wakes up and begins to hunt again.
The hole of marsupial digger includes main chamber located at the depth about
2 meters under ground, additional chamber which incorporates with main one
by unique vertical corridor, and system of radial burrows spreading from the
additional chamber to sides at the depth about meter. On the ground surface
it is possible to heaps of ground pulled outside by this animal. The marsupial
digger throws out ground through an inclined course, instead of vertical shaft,
as moles do.
Burrow systems of different animals of this species never incorporate directly
with each other: marsupial diggers are soliyary animals. But it yet does not
mean, that this animal avoids other “colleagues” in underground life: whenever
possible this animal builds joint systems of holes with rabbits. Some burrow
systems of marsupial diggers frequently adjoin from edges to one settlement
of rabbits. Obviously, rabbits are not in delight from such neighbourhood:
the marsupial digger makes it having especially gastronomic interests – it
can attack rabbit cubs, and at times attempts even healthy adult rabbit if
it will manage to cut off way to escape to prey. Besides having attached the
dwelling to hole of rabbits, the animal saves forces at burrowing of own holes,
chasing every possible inhabitant of rabbit holes. More often rodents – mice
and rats appear on its “lunch table”. Sometimes even other small predators
- marsupial hermins (Mustelogale
dolichocranus) become its prey, getting for hunting in holes of rabbits. Obviously,
at an attack against such rather dangerous animals the horn scute on muzzle
rescues the marsupial digger against wounds.
During breeding season males leave holes and wander in underground burrows
searching for females. They are extremely aggressive relatively to each other
and if two males will meet in hole, one of them can kill and to have eaten
another one.
At these animals constant pairs have not formed, and the destiny of posterity
is completely indifferent to the male. At any opportunity male even can eat
young animals if will to find a hole with cubs. The female gives rise only
to three – four underdeveloped cubs. They creep in pouch (opening back) and
grow in it about one month. After that the female leaves them in hole, and
regularly comes to feed. For one year the female has about three packs.
Three-month aged cubs already can freely move in underground holes. They leave
territory of the female and live independently. Within the first months of
life repugnatorial glands at them are not advanced, therefore their presence
can remain unnoticed by adult animals of this species. But up to the period
of puberty (at the age of about seven months) at young animals the smell appears,
and aggression sharply grows. The marsupial digger lives seldomly more than
four years.
Marsupial
spiketail (Spinurotherium leucospinus)
Order: Marsupials (Marsupialia)
Family: Spike-tailed marsupial rats (Spinurotheriidae)
Habitat: bush savannas of Australia.
In Neocene climatic conditions in Australia became more favorable for life.
Water had come to central areas of this island continent, having transformed
deserts to savanna with circuit of shallow-water rivers and lakes. In such
favorable conditions many new species of animals have appeared on continent.
The fauna of Australia of Neocene epoch represents fantastical combination
of descendants of native marsupial mammals and introduced by people placentary
ones. Epizooties of rabies and plague of carnivores have reduced chances of
survival of introduced placentary predators to zero, but the competition to
them had perfected features of adaptation at local marsupial carnivors, and
had permited to few survived ones to take place at top of food pyramid. Among
large-sized herbivores it is possible to meet not only graceful and harmonous
descendants of camels found due to the people new native land in Australia,
but massive marsupial animals too. And among small and medium-sized animals
marsupials successfully compete to rabbits and rodents introduced to this continent.
Among dense grass of Australian savanna it is possible to see strange object:
something similar to white flower, sticking up on hairy stalk. But, when this
object starts to move among grass stalks, it becomes clearly seening, that
it is not a plant but any animal. From time to time the owner of this “flower”
rears on hinder legs, and it can be seen more detailly. It is a cat-sized animal
covered with short wool. At it there are long flexible nose surrounded with
thin wiskers, round eyes and ears similar to rabbit’s ones. Well advanced claws
are appreciable on strong paws of this animal. And the thing looking like certain
fantastic “flower” actually is very long tail. At the tip of tail hairs are
modified to white-colored needles with black bases forming hairy brush. For
such appearance of this structure the animal had received the name “spiketail”.
It has habit to lift from time to time tail vertically upwards: this way spiketail
communicates with neighbours at long distance. Ancestor of the spiketail is
one of species of the Australian marsupial mice.
Colouring of animal is rather motley: the body is covered with brown wool with
dim black speckles, the mouth is led round by thin strip of white wool. On
the back of marsupial spiketail there are some alternating black and white
cross strips. All tail is black, and white needles are brightly appreciable
on its background. Being attacked by predator, spiketail warns it that it is
armed, curving back like a cat, lifting up tail, and shrilly squealing. If
it does not stop predator, the spiketail protects itself against it by impacts
of tail. In the basis of each spike there is a muscle attached by one end to
backbone. With the help of such muscles the spiketail can press spikes to each
other, forming “beater” or to spread them widely in sides, forming true weapon
which impact can be rather painful. Defending itself against predator, marsupial
spiketail turns around on hinder legs, keeping to aggressor by side and swinging
by tail extensively.
In mouth of spiketail it is possible to see both sharp incisors and canines,
and wide knobby molars. The marsupial spiketail has a little deviated from
diet of ancestors: it is omnivorous (marsupial mice are predators), and the
most part of its diet consists of plants. The animal is able to dig out roots
from ground, dexterously swarms up bushes and low trees in searches of fruits,
chews unripe seeds of grasses. But it also likes to diversify the diet with
food of animal origin. On trees and in bush marsupial spiketail searches for
small reptiles, large insects and bird’s nests. In the ground except for roots
spiketail willingly finds and swallows worm or larva of any insect. By smell
it finds the rests of predator meal, and with pleasure gnaws bones and cartilages.
These animals are solitaries, but they support visual contact to neighbours,
from time to time rising on hinder legs and looking around. They usually avoid
close meetings, preferring to leave an urine mark on objects allocated among
savanna – trunks of trees, termitaries and stones. Unique exception is a time
of breeding season. At this time marsupial spiketails gather to small groups
– approximately fifty-fifty females and males. Males compete with each other
for females, establishing hierarchy in short duels. During such tournaments
they beat each other by tail, but in this case needles are densely compressed
and do not cause wounds to the opponent. After pairing the male loses interest
to the female and starts to look after for next one. Thus it jumps after chosen
female on hinder legs, uttering keen chirp and snort. Each female couples with
several males, therefore paternity at these animals is difficult to establish.
Sometimes in one pack cubs at once from two males can appear.
After short pregnancy the female gives rise to about ten tiny underdeveloped
cubs. But the chance to survive is not at all of them: at the female there
are only six nipples. The pouch at this animal is presented by two longitudinal
plicas of skin, surrounding nipples from two sides. This skin is elastic, therefore
cubs are not threatened with danger to drop out of it. They develop in pouch
about three months, and to this time from six cubs only three or four ones
survive. Grown up cubs move on back of mother, and it drags them on herself
one month more. At this time on tails of young animals needles start to grow.
Gradually cubs pass from milk to adult diet, and start to feed near mother
independently, at the same time training at it to ways of food getting. Half-year
old cub already becomes completely independent. The young female can bear posterity
first time already at one-year-old age.
Marsupial
sloth (Bradyphalanger scandens)
Order: Marsupials (Marsupialia)
Family: Phalangers, or Climbing marsupials (Phalangeridae)
Habitat: tropical woods of Northern Australia and New Guinea.
In Neocene Australia moving to the north has collided against New Guinea Island.
As a result in northern area of continent young mountains have started to increase.
This area is in the zone of equatorial climate, and mountain covered by rainforest
giving shelters and food to various live creatures. As well as in other parts
of the world, life in Australian-New-Guinean rainforests is concentrated in
forest canopy at height 20 - 30 meters above the ground. Many original live
creatures, descendants of Holocene inhabitants of this region, live here. One
of them is the original herbivorous representative of marsupials – marsupial
sloth, the descendant of cuscuses (Phalanger).
Similarly to South-American “colleague” of Holocene epoch, marsupial sloth
is the inactive herbivorous animal slowly moving in tree crones, suspending
on them from below with the help of hooked claws. This animal weighs about
15 kgs; length of its body is about half meter. However, as against the “true”
sloths of South America, the Australian marsupial sloth has also long prehensile
tail, by the size exceeding body length. The bottom side of this tail is covered
with naked skin with set of cross combs (similar to papillar patterns on hands
of primates) – it raises durability of capture of support. Working as the “fifth
paw”, tail helps this animal to keep on branches. But basic organs of movement
are unusual paws of the marsupial sloth.
Forepaws of this animal are similar to monkey’s ones - the thumb opposes to
fingers. But on hinder legs from one side the thumb and joined together second
and third toes, and from other side fourth and fifth toes are opposed against
each other. Back legs of the marsupial sloth are powerful “pincers” by which
it grips branches when it takes to itself food by forepaws. Backbone and neck
of the marsupial sloth are very flexible: hanging a back downwards, the animal
easily can bend a half-revolution, touching by forepaws the bottom branch of
tree.
Head of the marsupial sloth is rather flat, short and wide: at it chewing muscles
are strongly advanced. Molars of animal are wide and knobby – leaves of trees
and firm seeds which this mammal cracks with loud crunch make significant part
of its diet. The marsupial sloth willingly adds food of animal origin to its
diet: eggs and nestlings of birds, and also large insects and snails.
The marsupial sloth is colored according to the habit of life: because more
than half of time it spends hanging under branches, its stomach is darker,
than back is. On the back of animal fur is gray-brown with numerous narrow
faltering cross strips – it is an excellent imitation of tree bark. On stomach
wool is monocoloured, dark-brown. Hands and feet of animal are naked, covered
with rosy-gray skin. Around of big orange-brown eyes there are circles of yellowish
wool, tip of muzzle is of the same color. Male and female do not differ in
colouring; the male is simply a little larger.
At this animal color binocular sight is advanced, there is sharp sense of smell,
and the hearing is a little bit dulled. Nasal cavity of the marsupial sloth
is short and wide, and ears are almost latent in fur. By the “intellectual
development” this animal continues “traditions” of marsupials: brain cavity
at it is small. But at its habit of life the special ingenuity is practically
not needed.
Marsupials sloths are sluggish solitary animals. They are practically omnivorous
and do not depend on any limited fodder resource, therefore they tolerantly
concern to presence of each other and have no protected territories. Only the
female bearing posterity in pouch becomes a little more aggressive to neighbours.
Pairing and birth of cubs at marsupial sloths occurs practically in any season.
The female ready to pairing, during movement marks branches and leaves by the
smell: repugnatorial glands at it are in axillary area, and it simply drags
a branch between paw and side, having griped it. The male finds the female
using these marks; then he couples to her without any rituals, and marks wool
of the female by his smell. Such feature of behaviour is characteristic for
some Holocene species of cuscuses.
Pregnancy lasts not a long time, and after 8 - 9 days tiny underdeveloped cubs
are born. Usually up to 10 cubs are born, but at the female there are only
six nipples, therefore some newborn marsupials sloths at once are doomed for
death. Cubs independently get into pouch opening back, and start to suck milk
and to grow. Approximately at the age of two months they can leave mother’s
pouch and remain on its body, clinging for its paws. Usually up to this moment
only three or four cubs from all survive, and only two or three cubs from litter
can become independent completely. At the age of four months youngsters already
depend on mother a little, and to the age of five months they become completely
independent. At this time the female already carries a new pack in pouch. For
one year it can give birth up to three packs of cubs.
Young marsupial sloths become sexual mature at one-year-old age though active
growth proceeds at them till two years. Life expectancy of these animals is
no more than 8 – 9 years.
Drummer cuscus (Onychowirlda tympanista)
Order: Marsupials (Marsupialia)
Family: Phalangers, or Climbing marsupials (Phalangeridae)
Habitat: plain rainforests and mountain fog woods in Northern Meganesia.
Neocene is the epoch of the occurrence of extensive large forests in Northern
Meganesia, and the development of rich fauna of tree-climbing animals. In forest
canopy, at height of several tens meters above the ground, various mammals
live: rodents, chiropters and marsupials. For similar ecosystems the combination
of two factors is typical: on the one hand, the abundance of kinds of food,
but on the other hand, rather rigid competition. Therefore in such ecosystems
very much specialized species frequently evolve.
In the evening and at night in forest canopy frequently it is possible to hear
rolling knock on tree, similar a little bit to drumming which a beak of woodpecker
beats out. Woodpeckers do not live in Meganesia, therefore instead of them
absolutely other animals hunt wood-boring insects. One of these creatures is
rather large (cat-sized) species of cuscuses. This creature is vaguely similar
to lemur aye-aye (Daubentonia), dwelt on Madagascar in Holocene epoch. But
instead of thin sensitive middle fingers thumbs armed with thick massive claw
are strongly advanced on forepaws of this cuscus. On other four fingers there
are very sensitive subungiuses covered with thin skin. Searching for insects,
this cuscus “drums” on bark and wood by thumb claws and catches vibrations
of wood by subungiuses of other fingers. Because of this feature the animal
has received the name drummer cuscus. Most likely, it is the descendant of
fox brush-tailed cuscus (Trichosurus vulpecula), one of few cuscuses had received
advantage from the neighbourhood with people in Holocene epoch.
This marsupial is fine tree-climber, spending all life in crones of trees and
never going down to the ground. All paws of this animal are prehensile, and
the long fluffy tail serves for maintenance of balance. The short velvety fur
of the drummer cuscus is colored grayish-beige color with marble pattern of
cross strips. It provides fine masking for animal swarming up on bark. Drummer
cuscus is too massive to jump from branch to branch similarly to marsupial
lemurs, cats or primates. But its paws differ in great strength of seize –
this animal can hang freely on one paw, and during the combat with small predator
cuscus can simply strangle it. Even dead animals of this species seize branches
so strong, that they fall on the ground already semi-decomposed.
At this marsupial there is the head with short wide muzzle and large sensitive
ears. Eyes of drummer cuscus are large and yellow, with strong shine characteristic
for nocturnal animals.
Distinctive feature of this animal is strongly advanced incisors similar to
incisors of rodents, and very strong chewing muscles. This is the adaptation
for feeding on insects hidden in thickness of wood. “Percussing” wood by claws,
animal listens to the tonality of sound, and feels by fingers changes in character
of vibration of wood. By these attributes drummer cuscus defines presence and
direction of tonnels made by larvae of insects in wood. Having found out a
tunnel pierced by beetle larva or other insect, cuscus opens it by several
strong bites.
Other feature of this mammal is very long tongue which can extend from a mouth
to ten centimeters. The basis of tongue is strongly shifted back - almost to
breast bone. On the tip of tongue the cross comb of corneous hooks, which can
be shifted relatively to each other a little, grows. Having opened dwelling
of insect, drummer cuscus pushes tongue in hole, and hooks larva on by these
hooks.
This animal goes to feeding in the evening and at night, and in dawn time it
hides in deep tree-trunk hollows or covered bird nests. Drummer cuscus dozes
hottest time of day, having covered sensitive eyes by ears and paws. At this
time it is enough vulnerable. Day time sleep of animal is sensitive, and, being
disturbed, drummer cuscus makes sharp movements by claws of forepaws, trying
to strike the enemy by them.
Drummer cuscuses are strictly solitary animals; only the female and her posterity
can keep together till some time. At night each animal from time to time reminds
to neighbours of its existence, uttering shrill high-frequency squeak. Male
searches for female ready to pairing, using such sound and special odorous
labels.
The male of drummer cuscus is approximately a quarter lighter, than the female
is; besides at him there is darker pattern on skin. Seasonal prevalence in
breeding at this animal is not expressed, and at any time of year in wood there
are females having posterity at various stages of development. Per one year
the female of this species gives rise two times to up to ten cubs, which leave
the brooding pouch at the 3-rd month of life. To this moment from pack only
two or three cubs survives. Till about one month they move in wood on back
of mother, and then start to lead independent life. Young animals reach the
sexual maturity at the second year of life. General life expectancy of animals
of this species does not exceed 10 years.
The idea about existence of this animal was stated by Simon, the forum member.
Ursine
cuscus (Ursiphalanger marsupialis)
Order: Marsupials (Marsupialia)
Family: Phalangers, or Climbing marsupials (Phalangeridae)
Habitat: New Zealand, flatland and mountain forests of temperate climate.
In different places of the Neocaenic Earth it is possible to find consequences
of human activity. Certainly, these species, as well as its many contemporaries,
have left the trace in paleontologic annals of planet. But there is also other
trace: results of introducing of various plant and animal species to other
habitats where they did not meet and where could not get naturally earlier.
The fauna of some places has very seriously suffered from rash installation
of new species, especially fauna of the remote islands.
When the human species had disappeared from Earth face, species delivered
by it at all did not gather to die out after it. Simply evolution in those
places where unbidden visitors have got has gone by other way, rather than
before. In fauna of different places species have appeared which never would
get there: in Australia various species of camels,
in Europe the descendant
of raccoon similar to bear have appeared, and isolated from all continents
New Zealands only due to activity of people had got ground mammal fauna.
One of the largest Neocaenic New Zealand mammals is the marsupial of huge sizes,
the ursine cuscus. It is the descendant of fox brush-tailed cuscus (Trichosurus
vulpecula), the marsupial mammal, acclimatized in New Zealand approximately
in 1900.
The New Zealand ursine cuscus is a huge ground animal weighting up to 300 kgs.
It is one of the largest representatives of marsupials of Neocene epoch. The
constitution of animal is massive, vaguely similar to ground sloths Megatherium
of prehistoric epoch.
The ursine cuscus spends the most part of life on the ground. The tail of its
ancestors, wood cuscuses, was any time adapted for branch seizing at climbing,
but at the ursine cuscus the tail is strongly reduced: it is short and nonflexible.
During walking of animal it serves as the balance weight, counterbalancing
the body. In the basis of tail the fat necessary for maintenance of ability
to live of an animal is accumulating – in winter the ursine cuscus becomes
languid and slightly hibernates This animal clambers on trees seldomly and
very clumsily, preferring to food on the ground. Its hind legs are plantigrade;
feet are wide, covered with thick layer of cornificate skin: it permits to
go easily both on stones heated up by the sun and on ice. The ursine cuscus
moves basically walking on two legs but frequently lowers to all four paws,
especially if it is feeding on mountain slope.
Wool of animal is thick, grey with black “cross” on back: longitudinal black
strip from nape up to middle of tail, and black stain on hips adjoining to
it. Throat, chest and stomach are covered with yellowish-white fur, and on
the end of tail the black hair brush grows. Due to thick warm wool animal can
normally live in mountains where in spring and an autumn there are light frosts
frequently.
At the ursine cuscus there is rather massive head with wide flat forehead.
In connection with nocturnal habit of life at it there are large eyes “shining”
in darkness as it is characteristic at cats. Pupils are vertical, iris of of
eyes is chartreuse. Ears are short and pointed. The ursine cuscus badly distinguishes
colors. But it perfectly sees in darkness, has good hearing and keen sense
of smell with which help correctly finds edible leaves and ripe fruits.
At this animal there are strong jaws, wide molars and large chewing muscles:
the significant part of its diet includes young bush sprouts, and also seeds
of grasses. Except for them the ursine cuscus eats roots and tubers of plants.
On forepaws of animal hooked claws serving for ground digging and defense against
predators grow. The thumb of forepaw is opposed to fingers – it is a heritage
of climbing ancestors appeared very useful, increasing dexterity of movements
at food getting. On thumb flat nail grows instead of claw.
The ursine cuscus has nocturnal habit of life hiding in bush thickets in the
afternoon. In the afternoon the animal is inactive, especially in hot weather.
At this time the ursine cuscus alternates feeding with the periods of short
deep dream. After day spending of animals there are traces of their activity
in bushes: broken off and picked branches, and also small holes in ground where
animals had slept. But at night animals leave shelter and wander in wood searching
for forage. They mark the route by odorous liquid from the special gland on
hip, sniff at marks left by other individuals. With the help of these marks
animals learn about health state and physiological condition of neighbours.
They do not avoid direct contacts to neighbours: neighbours treat peacefully
enough to each other, they are frequently feeding and have a day rest together.
The convinced singles among the ursine cuscuses are rarity. Usually ursine
cuscuses keep in groups of 3 - 4 adult animals, but structure of these groups
is changeable: in the same structure such groups exist maximally some days,
and then they break up - some animals leave it, other ones appear. Usually
each animal migrates on wide territory, but does not come out of its borders.
Borders of the “inhabited world” at different animals are not coinciding to
each other.
During winter cold snap ursine cuscus becomes sluggish and temporarily loses
desire to travel. Having found suitable place in wood or among rocks, the animal
digs out to itself shelter like den, or expands available one. In this shelter
the animal makes a litter, and hibernates. It not true hibernation: in this
condition the body temperature of the ursine cuscus is reduced only to some
degrees, it periodically moves and wakes up for a little time. At the ursine
cuscuses living on plains and in woods of the north of New Zealand, winter
lasts not for long – sometimes only about two weeks. At the animals living
in colder areas, it can be tightened about two months.
Pairing at these animals occurs in an autumn, but the embryo stops in development
up to the beginning of winter. In winter the female gives rise up to 3 cubs.
The pouch at this species opens forward. Cubs get into it (at this time the
female rummages hibernation for a little, cleaning pouch before cub birth)
and start to suck milk. They develop about 15 weeks, reaching to this time
of the size of kitten. At this age the pouch becomes cramped for them, and
they gradually move on back of mother. Keeping by paws for its wool, young
cuscuses gradually start to learn the world and to accustom to adult life.
Cubs try food eating by mother, study to distinguish other neighbours; sometimes
they even play. At the age of about one year, at weight about 60 kg, they begin
independent life. This is the most dangerous time for young growth: they not
always can expect for protection of adult animals if the predator will attack.
Growing up, young animals become capable to protect themselves. The main weapon
of the ursine cuscus is sharp claws on forepaws by which the animal inflicts
deep wounds to the aggressor.
At the age of four years young animals can bear posterity.
Marsupial grizzly (Gravipossum ursinus)
Order: Marsupials (Marsupialia)
Family: Predatory possums (Carnopossumidae)
Habitat: forests and woodlands of Meganesia, plains of southern part of continent
(up to coast of Eyre Gulf) and Great Dividing Ridge.
In Holocene Australian marsupials had suffered significant damage from the
introduction placentary predators to Australia, and from destruction of megafauna
by first aborigenes. At the continental part of Australia marsupial lions (Thylacoleo),
marsupial wolves (Thylacinus) and marsupial devils (Sarcophilus) had died out.
In historical time the Tasmania Island remained last shelter of marsupial wolf
(smaller, rather than its continental species) and marsupial devil (also smaller
than species exterminated at the continent). The place of marsupial predators
on continent had been actually occupied by people and feral dogs, dingoes.
In Neocene when both people, and placentary predators had died out of different
reasons, marsupials had started to evolve actively, occupying the exempted
ecological niches. Among new marsupial predators of Australia descendants of
both “true” predatory marsupials (Dasyuridae), and other animals passed to
predating in parallel to them had appeared. In Neocene possums had made the
second attempt to become carnivorous creatures (marsupial lion was the first
and rather successful attempt). Being not able to compete with dasyurids, carnivorous
possums had turned to massive animals, similar to bears by constitution. The
size of marsupial grizzly, largest one of them, reaches 3 meters in length
at weight about 500 kg.
Marsupial grizzly is one of largest species of marsupial mammals of the world.
Longteeth wombats,
descendants of Holocene wombats, reach even bigger size, but among possums
this species is true giant. Even huge New
Zealandian species of possum is much smaller.
Marsupial grizzly is very similar to massive bear with rather short extremities
and back curved upwards. At it there are plantigrade extremities armed with
sharp claws up to 10 cm long. Being attacked by predator, the marsupial grizzly
defends, rearing on hinder legs and delivering crushing blows by front ones.
It had not lost the ability to climb on trees though adult animals do it hardly.
But young marsupial grizzlies spend a lot of time in crones of trees where
they eat foliage and fruits, and ravage bird's nests. Adult animals feed mainly
on the ground.
Possums known to people differed in long tails. At marsupial grizzly which
spends on trees only small part of life, the tail became short and thick. It
has lost past function of the balance weight and the “fifth paw”, and now in
it the stock of fat is accumulated, helping to go through fodder shortage.
Head at this marsupial is rather large and massive. Ears of animal are short
and round; nose is big and lips are mobile. Eyes of marsupial grizzly are small
– the animal is short-sighted, and in search of food bases at sharp sense of
smell. Marsupial grizzly has yellowish-grey fur with dark paws and nose bridge.
Stomach of animal is white, on back from waist up to the basis of tail narrow
black strip stretches. Feet and hands of animal are covered with thick cornificate
skin.
Jaws of marsupial grizzly are short and have slightly increased canines and
wide molars. Strong chewing muscles help this marsupial to chew fibrous parts
of plants and heads of bones of large animals. This species of animals partly
plays a role of scavenger in ecosystems of southern and eastern parts of Meganesia.
It finds rests of catch of large local predators, and eats them almost completely:
after the feast of marsupial grizzly only separate vertebrae and bones of legs
remain from carcass of animal. Also it can drive away smaller predators from
catch. The males of this species not burdened with cubs bearing can go fishing
in shallow rivers and lakes.
Marsupial grizzly is not completely carnivorous species of marsupials: it is
omnivore, and whenever possible willingly eats vegetative food. With the help
of long claws animal digs out roots of ferns and other eatable plants. It can
dig out holes of small animals just as it was done by bears in Holocene epoch.
Because this animal is omnivorous, at it the increased intraspecific aggression
characteristic for carnivorous species is not shown. Besides females at marsupial
grizzly are larger, rather than males, therefore cubs are not threatened with
danger of attack from their side while they stay with mother.
Brooding pouch at this species of mammals opens back, and there are four nipples
in it. Fertility of the female of marsupial grizzly reaches ten cubs, but only
four of them receive an opportunity to survive, and usually two cubs survive
up to independence. In the beginning the posterity of marsupial grizzly is
“worn” in mother’s brooding pouch over half-year. After that cubs change to
back of mother, and keep by paws for her wool. The posterity stays with mother
up to one-year-old age, and right when young animals abandon the female, it
becomes ready to pairing again.
At the eastern slope of Great Dividing Ridge, in humid mountain woods the relative
of this species, the marsupial panda (Gravipossum paraailuropoda) lives. This
species is much smaller, rather than marsupial grizzly: the adult female weighs
about 300 kg at length of body up to 2 meters, and the male is 25 – 30 kg lighter.
In food predilections marsupial panda differs from marsupial grizzly by appreciable
bias in vegetarianism. The food of animal origin amounts a small part of diet
– marsupial pandas eat eggs of ground-nesting birds, and willingly consume
carrion. Frequently animals search for forage on banks of mountain rivers,
feeding on water invertebrates and local migrating fishes of Galaxiiformes
order. But the basic food of this animal is made of roots and leaves of grassy
plants, and also sprouts of bushes. Living in cool mountain climate, marsupial
panda differs in rich wool. At this animal there are black head, shoulders
and back, rounded white spots above eyes, grey sides and back part of body.
The tail of marsupial panda has snow-white inner side which is used by animals
for submission of signals to congeners.
Fertility of marsupial panda is insignificant: the female gives rise to no
more than five cubs from which only one or two ones survive up to independence.
Life expectancy of this animal reaches 40 years.
Marsupial biruang (Carnopossum heliarctiodes)
Order: Marsupials (Marsupialia)
Family: Predatory possums (Carnopossumidae)
Habitat: mountain and lowland rainforests of north and northeast of Meganesia.
In Neocene Australia and New Guinea had merged to unite land massive named Meganesia.
This continent continuously moves to the north, to the equator, “crushing” edges
of lithosperic plates of Indonesia. Actually, this circumstance became the reason
of raising of northern part of Meganesian lithosperic plate.
Movement of continent has caused the considerable changes in climate of territories
included in continent. The northern Meganesia in Neocene is in the area of equatorial
climate that has positively had an effect on nature of continent. The north is
covered with rich rainforests. And in forest swamps rivers originate and flow
to the south and carry water to areas represented rigorous waterless deserts
in Holocene.
Rainforests of the north of Meganesia are rich in life. Various birds, reptiles
and amphibians, and also unnumerable species of insects live here. Besides, the
forest canopy is inhabited by various small and medium-sized mammals, and large
species of mammals are found in underbrush. Among them there are descendants
both of native Australian and New Guinean species, and descendants of species
introduced by people.
Rainforest is a habitat of rather large species of marsupial mammal which is
named marsupial biruang. It belongs to group of carnivorous possums classified
in family Carnopossumidae. Its close relatives are huge marsupial grizzly living
in the south of continent, and marsupial panda – smaller species from mountain
areas at the east of the continent. Marsupial biruang is the smallest representative
of carnopossumids: weight of adult animal is only about 60 kg. It is the ecological
analogue of Malayan bear, or biruang, known in human epoch – hence the name of
this species.
Marsupial biruang has robust constitution and rather short limbs. Similarly to
all carnopossums, it has very short tail which does not take part in tree-climbing.
On paws of animal long hooked claws grow; with their help marsupial biruang easily
gets on high trees. Fingers of animal are mobile and permit it to gather very
small objects and to catch insects. The wool of this animal has black or dark
brown color, and only above eyes there are two light grey spots. At young animals
spots above eyes are absent. On hands and feet wool is thin, and through it grey
skin of animal is visible.
Eyes of animal are directed forward, and provide enough wide sector of three-dimensional
vision. Ears at marsupial biruang are short, rounded, slightly jutting out from
wool. At this animal there are rather short muzzle and increased forward cutters.
Molars are obtused: their shape is characteristic for omnivores.
Marsupial biruang uses for life all levels of rainforest – from underbrush up
to forest canopy at height over 20 meters. This is sluggish single animal marking
borders of territory with the help of musky liquid secreted by two glands at
the root of tail.
This animal is omnivorous and easily finds enough of food. It eats fruits of
various tropical trees, sappy leaves and stalks of epiphytic plants. Also this
animal catches large insects and digs out of ground worms and grubs. If the opportunity
is represented, marsupial biruang catches little mammals and ravages nests of
birds. Any animals from insects and snails up to vertebrates weighting up to
3 – 4 kgs become its prey.
Having caught prey, marsupial biruang perches on wide strong branch of tree,
sits on hinder legs and eats food, holding it in forepaws. After meal the animal
licks wool by long tongue, and “washes” by forepaws like cat. Marsupial biruang
is active in day time, and only in hottest time of day sleeps till some hours,
having chosen a shady place.
At this species of marsupials the female is larger and heavier, than the male.
Animals do not form constant pairs, and meet only for the period of pairing.
The pouch at the female opens forward – when female swarms up a tree, such position
of pouch is safer for the large cub. The female bears only 2 – 3 rather large
cubs (large to measures of marsupials – newborn marsupial biruang weighs about
10 grammes). Some months later only one cub from the litter usually survives.
The female looks after it for a long time: the cub stays in pouch till about
seven months. Having left the pouch, it follows the female and learns to search
for food till about three – four months. At this time the female can couple with
another male and bear new posterity. When the new cub is born and gets into a
bag, the female starts to show aggression to the previous cub, and it passes
to independent life.
Sexual maturity at marsupial biruang comes at the age of four years, and life
expectancy reaches 40 years and more.
The idea about existence of this species of animals was proposed by Simon, the forum member.
Night
marsupial lemur (Nyctiphalangeropsis strigops)
Order: Marsupials (Marsupialia)
Family: Marsupials lemurs (Phalangeropsidae)
Habitat: rain tropical woods of Meganesia.
The canopy of tropical rainforests stretched at height 20 - 40 meters gives
a plenty of opportunities for adaptation to various live creatures. Among its
inhabitants it is a lot of marsupial mammals. Those species have survived at
the border of Holocene and Neocene, have inherited from missed species many
free ecological niches, and have occupied them, having evolved to fantastical
lifeforms. The competition between inhabitants of rainforest canopy can be
very rigid, therefore during evolution live creatures developed to lower it
as much as possible. Certainly, it is possible to become the specialized “expert”
in consumption of the certain kindss of forage, and it is possible simply to
live and not to prevent to another ones. One of animals living in this wood
had made it.
When night falls on wood, the majority of wood inhabitants hides in tree-trunk
hollows and nests and falls asleep. But some animals just at this time wake
up and quicken. Among them there are various insects, frogs, lizards, some
birds, and also small mammals covered by black wool with white marks. Their
movements are very similar to movements of monkeys; their heads are big, have
high cheek-bones and large “owline” eyes. Animals keep in small group, supporting
contact with the help of quiet chirp. They are similar to small monkeys, but
they are not primates at all, but descendants of cuscuses – night
marsupial lemurs.
The night marsupial lemur is insignificant: length of its body is about 20
cm, tail length – about 30 cm. All paws of these small mammals are adapted
to branch gripping: on forepaw there is well advanced thumb, and toes on hinder
legs are opposed to each other by two groups: joined together I, II and III
against joined IV and V. Bent claws grow on ends of fingers and toes. The animal
moves on trees like squirrel: it runs on branches and swarms up trunks, clinging
by claws against bark. Due to tenacious paws night marsupial the lemur dexterously
swarms up the thinest branches. It is able to jump from tree to tree like monkey:
having shaken on thin elastic branch, the small mammal makes jump some meters
long. During flight in air the mammal stretches paws in sides to glide a little
bit and thus to increase range of jump. The prehensile tail increases opportunities
of animal: due to it night marsupial lemur can hang on branches.
The nocturnal habit of life is reflected at the appearance of this animal:
eyes of the night marsupial lemur are very large, and because of it the obverse
part of skull looks rather wide. On muzzle of mammal long whiskers grow with
which help the animal finds way in dark night wood. Colouring of wool is coal-black,
but for recognition of neighbours at night marsupial lemurs there are contrast
marks – white stains on chest and tail tip. Around of animal’s eyes there are
rings of white wool because of what this small mammal looks a little horribly:
rings create illusion of eyes belonging to larger animal. Eyes of this marsupial
lemur are yellowish-brown; pupils are vertical, as at cats. This animal distinguishes
colors badly, but it is capable to see world around even at weak light of stars
in moonless night.
At night marsupial lemurs there is good hearing, though auricles very short.
But the ear aperture at them looks like a crack, and on back edge it is edged
by fringe of rigid fur functionally replacing an auricle. Due to sharp hearing
the animal can search for insects in darkness, and frequently to it succeeds
even to hear flying owl: it is rather useful property for nocturnal animals
of small and medium size.
The night marsupial lemur is a sociable animal; it lives in groups of 3 – 5
individuals. All group consists of male and several females, and also of cubs
have not reached independence. These animals spend day in tree-trunk hollow
on their territory. They know an arrangement of all suitable shelters on their
area, and each morning they stay in new one. By this way animals prevent the
parasite breeding and hide their resting place from predators.
Night marsupial lemurs wake up, when day time heat falls down, and day time
predators search shelter to lodge for a night. Usually dominant male or the
main female shows itself first from tree-trunk hollow. The animal cautiously
smells air, convincing of safety, and then comes out on the branch. After it
other members of group appear. Animals clean wool, stretch, and start to inspect
territory. They move on branches in a line, keeping a distance. During the
night small mammals become especially cautious: at this time owls fly to hunt.
To protect itself against attacks of these birds, night marsupial lemurs choose
way in richest branches, and jump from tree to tree one by one, stopping for
a little time on trunk after landing. Other enemies of these animals are snakes.
The silent rustle similar to sound of the moving snake is the most terrible
sound for night marsupial lemurs.
The group marks borders of the territory with the help of secretions of repugnatorial
glands located in armpits. For this purpose they simply drag branch between
body and paw. All members of group mark territory in same place, but on different
branches, therefore smells of separate animals do not mix up also any newcomer
can estimate number of animals protecting this territory. However, dominating
male will do not be against at all if the newcomer will appear the female.
When it is a lot of forage, it can admit to clan even other male, but already
beforehand a role of satellite and “whipping boy” will be determined to it,
and it will druggle at the end of group. But the solitary animal in almost
cases is restrained with such role: it is easier to notice danger together.
During the movement group supports contact: animals utter chirp and abrupt
sounds similar to bark.
Night marsupial lemurs are omnivorous: they will ravage the bird's nest or
will tear off ripe fruits from branch with the same pleasure. In searches of
vegetative forage they are guided basically by sense of smell, correctly distinguishing
ripest fruits by smell. One more dainty is thick juicy stalks of some orchids.
The best food of animal origin includes insects and spiders. If fruit appeared
worm-eaten, these mammals will to have eaten it entirely, right including caterpillar.
The special dainty is delicate soft termites. At night these blind moisture-loving
creatures leave their shelters, and night marsupial lemurs willingly lick them
off from bark.
Seasonal prevalence in breeding at night marsupial lemurs is not present: practically
at any time it is possible to find groups with cubs. Female twice per one year
gives rise to 7 - 8 cubs though it can feed only four ones. The pouch opens
back at this species. Cubs spend in it about two months, then they get out
on back of mother and travel on it, seizing its wool. This time is the most
important for their further life: cubs study to search for food and seize skills
of tree-climber. When they wean from dairy diet, their connection with mother
weakens. Cubs can get on backs to other members of group though those can be
not so pleased with it, and seldomly bear too long driving another's “youngs”
on the back.
Sooner or later young animals abandon clan. They go to “neutral” (“drawn”)
boundary territories of different clans, and gradually teenage groups break
up. Young females more often succeed to join to other clan, and the destiny
of males develops more dramatically: at them probability to finish the barchelor
life in stomach of the snake or an owl is most of all. Occasionally the young
male finds teenage group and expels from it other males, forming a high-grade
clan. Being lucky it can “overthrow” the old male and take hold of available
clan.
Life expectancy of the night marsupial lemur is short: it seldomly lasts more
than 10 years.
Hook-fingered kangaroo (Gravimacropus macrocheirus)
Order Marsupials (Marsupialia)
Family Kangaroos (Macropodidae)
Habitat: light forests and bush thickets of Southern, Central and Eastern
Australia, foothills of Great Dividing ridge.
During the evolutionary history kangaroes belonged to among the most successful
marsupial animals of Australia. They evolved rather quickly, adapting to
changes of world around, and in historical epoch competed on equal terms
with sheeps
and camels introduced by people. In Neocene the significant part of domestic
animals introduced by people to Australia, had died out, and only camels
evolved to some species adapted to various habitats became real competitors
of kangaroes.
Kangaroes had also evolved, and among them various kinds had appeared: some
of them eat foliage, others graze, and some ones even eat insects and carrion.
Climatic conditions of Neocenic Australia favour to development of large
life forms, and among kangaroes one such species had appeared.
The largest kangaroo of Australia is the hook-fingered kangaroo. It is huge
species up to 3 meters in height in biped pose. The hook-fingered kangaroo
has shape characteristic for representatives of this family, but adjusted
for larger size and massive constitution. This animal weighs about 250 -
300 kg,
therefore it jumps slowly and hardly, more preferring to move by original
slowed down “gallop”, moving alternately pairs of front and hind legs. Forepaws
of
the hook-fingered kangaroo are adapted to such type of movement: the animal
leans on the external side of long hands covered with thick skin, similarly
to gorilla. On fingers of this animal thick hooked claws grow. At movement
the animal pushes from the ground by tail which is considerably short in
comparison with tails of usual species of kangaroes. On the bottom side of
tail the cornificated
covering, similar to callouses developing on legs of camels, is advanced.
But this “callous” does not serve for protection against burns as at camels,
but
provides the best cohesion with ground at the movement.
Hinder legs of the hook-fingered kangaroo are lengthened: it is a characteristic
attribute of all kangaroes. But they are not much longer than forepaws, short
of a foot. Foot at this species of kangaroo is long and elastic, and claws
are not sharp, but to a greater extent hoof-like and blunt. The hook-fingered
kangaroo does not concern to list of fast runners: it prefers to not escape
from predators in flight similarly to the majority of its congeners, and
to protect itself in close combat. This animal defends against enemies by
impacts
of forepaws, gashing it by terrible lateral impact strengthened with a turn
of all body. This animal counter-attacks large enemies and tries to tear
them by claws of hinder legs.
The wool of hook-fingered kangaroo is colored bright - red. Head and forepaws
at it are dark, almost black, and hinder legs and the bottom part of tail
are lighter, than the basic colouring of body is. On head there is the extensive
white “mask” covering muzzle, eyes and a part of bottom jaw.
This animal is original analogue nowadays extinct sharp-clawed perissodactyls
chalicotheres (Chalicotheriidae). The hook-fingered kangaroo eats leaves
of bushes and trees. Usually it is feeding, standing on hinder legs and tightening
tree branch to a mouth by long forepaws. Also it eats soft species of grasses,
not competing with other species of kangaroes eating rigid siliceous graminoids.
Chewing muscles are advanced rather poorly at it: the animal chews a little,
and the food is processed mainly in the complex stomach consisting of three
chambers. Symbiotic protozoans improving quality of food digesting, live
in stomach of this species. The feature of physiology of the hook-fingered
kangaroo
is the ability to make some of metabolic water which is formed at fat oxidation.
This adaptation helps this animal to live long time without water: the hook-fingered
kangaroo comes to the watering place only once a week, and even less often.
Jaws of hook-fingered kangaroo are short, and the obverse part of skull is
short and expanded. Eyes are shifted in sides and provide the good circular
field of view. Also at the animal there are wide mobile ears: hearing at
the hook-fingered kangaroo is keen. The olfactory cavity also is expanded:
therefore
the head of animal has no “deer” shape usually characteristic for kangaroes.
Wide ears and nasal cavities help large animal to cool itself in strong heat.
This species of kangaroo is solitary animal though it does not avoid a company
of congeners. In places rich in forage this species meets by small groups
of 4 - 5 adult animals. Usually animals keep together during pairing. The
courtship
season at this species is strongly dragged out, and females ready to pairing
meet the most part of year, except for the droughtiest months. The female
ready to get posterity, emits the special smell on which it is found by males.
Usually
one female is looked after with true “retinue” of males. But one of them
shows the greatest activity, driving other ones away. Fights between males
are limited
to struggle with the help of forepaws. Kicks in stomach of the contender,
characteristic for other kangaroes, at this species are excluded: animals
are too massive
for it.
After short pregnancy the female gives rise to one small cub, which should
get into parental pouch itself similarly to posterity of all marsupials.
The period of bearing of posterity in pouch at the hook-fingered kangaroo
is very
long: about half-year the cub constantly sits in pouch, sucking milk, and
still almost half-year it is hidden in it at the case of danger. But up to
age of
fifteen months the cub continues to suck milk of mother though to another
nipple its younger brother has already attached. Young animals differ from
adults
in monotonous colouring of body. It changes approximately to three-year age,
and four-year-old animals already can breed. Life expectancy at this species
may be over forty years.
The idea of existence of this animal was proposed by Simon, the member of forum.
Monkey wallaby (Pithecomacropus agilis)
Order: Marsupials (Marsupialia)
Family: Kangaroos (Macropodidae)
Habitat: rainforest of Hawaiian islands (Oahu, Molokai, Lanai, Maui, Kahulawi),
forest canopy.
Kangaroos never lived on Hawaiian Islands, similarly to representatives of
other groups of ground and flightless mammal. But in human epoch they have
appeared even in this isolated corner of Earth, being introduced there
by people. And it was species of mountain wallabies Petrogale penicillata (brush-tailed
wallaby) became extremely rare in true native land, in Australia, later. Pair
of these animals has run away from a menagerie in the beginning of XX century,
and was multiplied on Oahu Island. Decrease of ocean level till an ice age
had permitted to this animal to settled on the near islands: Molokai, Lanai,
Maui and Kahulawi. Obviously, this moving was unitary: Oahu Island is separated
from them by rather deep Kaiwi passage which had kept in ice age, and obviously,
it had been crossed casually by a little group of animals – founders of new
population. And the further moving of wallabies to the east on islands of the
Hawaiian ridge was blocked with Alenuihaha passage separating these islands
from Hawaii Island. Therefore moving of descendants of marsupial immigrants
on islands is limited to these islands. In the period when the volcanic island
Hekeua had been formed, the part of populations of this kangaroos has almost
disappeared, having kept in small amount only in rests of woods at coast. But
later, due to characteristic for marsupials ability to fast breeding, the population
of Hawaiian wallabies was restored. So these marsupials relieved by the nature
of competitors, others large mammals, have remained on Hawaii.
The monkey wallaby has considerably changed a habit of life characteristic
for ancestors. Now this is a forest animal, something similar to the “true”
Holocene tree kangaroos of New Guinea. It is the small species of kangaroos:
it is cat-sized animal, but seems larger because of long tail. This animal
has found a fine ecological niche in woods and became the largest species of
herbivorous animals in rich canopy of the Hawaiian rainforest.
Monkey wallaby dexterously swarms up trees. Joints of back extremities at it
are very flexible, and the foot is partly swivels sideways: the animal clasps
so a tree trunk more densely. On front and hinder legs hooked claws are advanced.
Grown together toes on hinder legs serve for the toilet purposes. This animal
clambers on trees “galloping”, working synchronously in pairs by forward and
back extremities. Perhaps, the monkey wallaby is unique species among kangaroos,
able to move back: this animal goes down from trees a tail first, serially
grasping a trunk by front and hind legs. Similarly to monkeys, this animal
can make big jumps from tree to tree, and is able to swarm up thin lianas dexterously.
If the branch or liana is especially thin, animal is able to move on it similarly
to sloth - a back downwards, clinging for a support by bent claws. A tail of
this species is thin and long: in length it exceeds body and head taken together.
Tail is not prehensile, but elastic: it serves as the balance weight during
jumps and movement among branches.
The skull of monkey wallaby is rather short and wide, as gives to this marsupial
the “monkey” shape. It is connected to several features of its habit of life.
First, the important advantage to this species is the binocular sight assisting
to estimate distance up to the next branch precisely. And second, the animal
eats rather soft food: fruits and soft leaflets, therefore its jaws became
weaker and have decreased in size.
Sight of the monkey wallaby has one feature: the animal can perceive ultra-violet
light. Therefore animals of these species easily find among foliage green fruits
covered with wax, reflecting this part of solar spectrum. Ears of the monkey
wallaby are wide and mobile; animal has very good hearing. Ears are hairless,
pink due to blood vessels appearing through skin: in hot weather vessels extend,
and surplus of heat is emitting through ears.
Coloring of body of monkey wallaby is disruptive: head is black with longitudinal
white strips stretching from nose to ear through eye, on forepaws and shoulders
there is light red “cloak”, tail is grey with black tip. Waist and hinder legs
are dark brown.
Being similar to monkeys externally, this animal is absolutely not similar
to them by habits: monkey wallaby is solitary animal, and two adult animals
can be together only in short courtship season. Monkey wallabies are very territorial
animals. Each individual occupies territory about a quarter of square kilometer,
and marks its borders by urine. This animal does not arrange constants shelters,
and spends night in casual refuges. And adult strong animals spend the night
even simply on branches.
Two times per one year the female brings posterity: one underdeveloped cub
growing in pouch similarly to other species of marsupials. The three-monthly
cub starts to get out of pouch, and at the age of four months it leaves mother’s
pouch finally. About one month the young animal studies at mother in searches
of forage, and then in courtship season she banishes the posterity. Young monkey
wallabies begin able to breed at two-year-old age.
Ostrich kangaroo (Theriostruthio unidactylus)
Order: Marsupials (Marsupialia)
Family: Kangaroos (Macropodidae)
Habitat: grasslands and semideserts of Meganesia.
Right after disappearances of mankind Australia represented the continent the
most part of which had been occupied with deserts and poor savanna. In Neocene
when Australia had merged with New Guinea to unite continent Meganesia, the
climate promoted the occurrence of productive savanna. This area is inhabited
by large flightless birds and descendants of camels introduced to Australia
in ancient historic time, similar to giraffes and antelopes. But representatives
of indigenous Australian fauna, marsupial mammals, successfully compete to
them. Among them the original species became ecological analogues of antelopes
and other quickly running herbivores had evolved.
The extreme degree of specialization to such habit of life is demonstrated
by huge descendants of medium-sized wallabies of Holocene epoch, representatives
of genus Theriostruthio (literally: “ostrich beast”) for which strongly expressed
gracile constitution and ability to high-speed moving on a long distance are
characteristic. These animals are a vivid example of convergence with running
birds and fossil ostrich dinosaurs (Ornithomimidae).
Ostrich kangaroo is settled at the significant part of Meganesia where savannas
were stretched. This animal lives in big herds numbering up to 100 individuals.
These animals are constantly in movement, and continuously migrate, not staying
at one place for a long time. Only in rain season when the grass plentifully
grows, animals do not hurry up to leave rendered habitable places. It is very
large species of marsupials – growth of the adult individual standing on rear
legs reaches 3 meters, and the length including the tail is up to 4,5 meters.
Ostrich kangaroos strongly differ from species known in human epoch. At them
there are very long and rather thin extremities. On hind legs of these animals
there is only one middle toe with wide blunt claw similar to horse hoof. It
is an attribute of quickly moving animal: usual speed of ostrich kangaroo makes
45 – 50 kms per hour, and animals pursued by predator accelerate momentum up
to 80 - 90 kms per hour. These animals perfectly move on firm ground, but are
compelled to avoid areas with soft and sandy ground.
Forepaws of this animal are approximately one third shorter, than back ones,
and on them only three fingers with thick dulled claws remained. They are mobile
enough to permit the animal to make some actions with their help. With the
help of these fingers the female of ostrich kangaroo clears the pouch before
the cub birth. Also ostrich kangaroos are able to break off tops of grasses
by forepaws and to eat them, holding body in vertical position. Such pose is
very characteristic for these animals: from height of the growth ostrich kangaroos
easily notice predators.
Neck of this animal is longer and more mobile, than at usual kangaroos. If
grass is short and thin, ostrich kangaroos can graze, moving on four limbs
by slow “gallop” characteristic for kangaroos (ostrich kangaroos are not able
to walk). Thus they graze grass by mouth.
The diet of ostrich kangaroos includes rigid and dry grasses, and also plants
which can be inedible for the camelopes. In connection with such food jaws
of ostrich kangaroos are short (their muzzle is similar to muzzle of large
fossil kangaroo Procoptodon), and teeth have folded enamel and are adapted
for masticating of highly abrasive food. Besides they accrue for rather long
time, before their roots will be formed finally. It helps to compensate deterioration
as a result of feeding on rigid vegetative food. Muzzle of ostrich kangaroos
is wide and short, with strong chewing muscles. Its outlines are especially
emphasized by large auricles. Such auricles are necessary for more effective
heat emitting – ostrich kangaroos inhabit hot and dry areas of Meganesia. They
can not drink for a long time, being content with metabolic water which turns
out at oxidation of food and fat.
Wool of ostrich kangaroos is short and velvety, colored in light shades – at
different individuals it varies from yellow up to almost snow-white and light
grey. Light color of wool rescues animals from overheating – in places of their
inhabiting it is almost impossible to find trees giving enough shadow.
Each year at the female of ostrich kangaroo only one cub is born. It comes
into the world small and underdeveloped, and independently creeps to the pouch
of mother. In pouch it attaches by mouth to the nipple and within several months
almost continuously sucks milk. The eight-monthly cub ventures to leave the
pouch for the first time, but at first attribute of danger it hides in it again
though it is rather difficult for it to make it with its long legs. Gradually
young animal starts to explore the world around, and passes to adult diet.
At the age of fourteen months the young animal finally abandons the parent
pouch and does not suck milk any more. The three-year animal already almost
grows till the size of adult individuals, and at the age of five years ostrich
kangaroos become able to breeding. Life expectancy of this animal reaches 40
years and more.
In territory of Meganesia more some species of ostrich kangaroos still:
Dwarf ostrich kangaroo (Theriostruthio nanus) lives in the strip of dry savannas
and semideserts to the west of Eyre Gulf. Its length does not exceed 150 cm
including the tail, and growth of sitting animal is no higher than one meter.
On the body of this animal there is a pale pattern of dense vertical strips
which pass to more contrast cross-striped pattern on tail and rear legs. Such
colouring of wool appears fine masking at life in high grass. This species
differs in high speed – at the short distances dwarf ostrich kangaroo can reach
the speed of more than 80 kms per hour. The female of this species can bring
posterity two times per one year – in spring and autumn. This animal grazes
on four legs only in thin and short grass. Usually dwarf ostrich kangaroos
eat, biting off tops of high grasses during the movement.
Striped ostrich kangaroo (Theriostruthio zebratus) lives in rather damp district
– in bushy savanna at the western coast of Eyre Gulf. It differs from common
ostrich kangaroo in considerably smaller size (length of body is up to 3 meters)
and darker colouring. On yellowish-brown skin of this animal brown vertical
strips stretch. Due to such colouring grazed animals are poorly appreciable
among grass and in light forest. These animals have also special feature of
behaviour – during movement each animal from time to time makes almost vertical
jumps of height up to 4 meters.
Forest ostrich kangaroo (Theriostruthio sylvestris) has the same size, as the
previous species. It lives in light forests growing in river valleys at the
east of Meganesia. Jaws of this animal are weaker and lengthened more, than
at common ostrich kangaroo. It is connected to a diet of this animal – it eats
leaves of undersized trees and bushes, and also large grasses. The body of
this kangaroo has spotty colouring – reddish-brown with vertical white strips
on back and shoulders. Also the throat and bottom part of head have white colouring.
Ears of this animal are rounded and have smaller size, rather than at species
living in plain district.
Mirriuula (Diablowallabia camelophoneus)
Order: Marsupials (Marsupialia)
Family: Kangaroos (Macropodidae)
Habitat: Meganesia, mountains in the east, to the south up to Flinders Range;
forests and bush.
The majority of kangaroo species represents herbivores of plains, bush and
mountains. But it is not only ecological niche which these animals could occupy.
The species of carnivorous kangaroo rats Ekaltadeta ima weight about 15 kgs,
and also one more predatory kangaroo, Propleopus oscillans lived in the past
in territory of Australia. In human epoch some wallabies differed in propensity
to zoophagy. After extinction of placentary predators because of epizooties
various species of marsupials occupied an ecological niche of carnivores during
Neocene with different degrees of success. The small naked-breasted kangaroo,
scavenger kangaroo rat (Harpotorous
microraptor), lives in semideserts of Meganesia,
eating carrion and the rests of prey of large
predatory reptiles. The result
of other attempt of kangaroos to develop predatoriness is mirriuula, the descendant
of mountain wallaby (Petrogale) living in mountain areas of Meganesia.
Mirriuula is named after mythical demonic dog from folklore of Australian aborigenes,
and quite justifies this name. It is a furious kangaroo weighting about 100
kgs, with flexible torso and long fluffy tail serving for balance. It is a
predator eating birds, herbivorous kangaroos and mountain
camelopes.
Mirriuula is not too similar to usual kangaroos by its constitution. It has
rather short hind legs, and rather seldom rears on them. The mode of its movement
is galloping jumping and true gallop. Mirriuula moves only on hind legs very
seldom – usually it does it, attacking prey. At this time its forepaws are
free, and mirriuula seizes prey by them. Mirriuula can run quickly both on
short, and on long distances, but on open district or in foothills it frequently
conflicts with other marsupial predators. But in mountains this species is
out of competition. It gets big advantage in hilly terrain over marsupial jaguar
and marsupial panther, surpassing them in dexterity. Besides these predators
are more heat-loving and prefer not to come into mountains. And mirriuula is
able to chase prey on hillsides; it arranges ambushes on tracks and frequently
hunts in mountain forests and among thickets of undersized eucalypti. Colouring
of wool of the mirriuula helps it to remain unnoticed. The basic background
of colouring is brownish-grey, darker on head and paws. Tail is light grey,
and its tip is almost white. On back of animal there are some longitudinal
lines of dark spots.
Having long and strong limbs and hard bent claws, mirriuula can easily catch
up and kill the goat-sized prey. It snatches on prey and brings it down on
the ground with the help of strong forepaws, and then tears its throat by long
pointed incisors slightly directed forward and closed, as a nipper. With the
help of incisors this predator can easily bite through neck of prey. In strong
jaws cutting molars with jagged edge and peaked molars easily cracking bones
and tearing meat from them grow.
The muzzle of animal resembles cat's one a little – it is short and high, with
large mobile ears. In nasal cavity of animal the olfactory epithelium forms
many plicas; therefore sense of smell of this animal is very keen.
This is solitary animal strictly protecting its territory. At the territory
of mirriuula there are some shelters, one of which is used constantly, and
others are temporary; animal spends night in them, when eats large prey left
near them.
At the territory of Meganesia smaller relative of mirriuula lives – fox
wallaby (Diablowallabia omnivorodentus). This species inhabits plains overgrown with
high grass, where it builds shelter representing deep hole. It hunts or traps
in ambush small vertebrates, eats insects, digs out grubs, nutritious stalks
and tubers of plants. It can run very quickly – as quickly, as its large relative.
This animal has larger ears resembling ears of fennec fox – it is the adaptation
for heat emitting. The wool of fox wallaby is colored yellowish-grey with several
longitudinal lines of brown spots.
These species of animals were discovered by Tim Morris, Adelaide, Australia.
Digging armadillo (Postpichisiego trachycephalus)
Order: Edentates (Edentata)
Family: Digging armadilloes (Armotalpidae)
Habitat: South America, pampas and light forests from warm temperate up to
tropical zone.
In human epoch in South America mammals constantly leading digging habit of
life (the analogues of moles of Holarctic region) were absent. At the same
time in territory of this continent there were many kinds of burrowing rodents
and armadilloes. Fairy armadillo (Chlamydophorus truncatus), or pichisiego
died out in human epoch had led the way of life closest to moles.
In Neocene of South America armadilloes had taken a significant place in fauna.
They successfully compete to rodents, and one species became a
large scavenger.
Among Neocene armadilloes one species which conducts practically completely
underground way of life had evolved. This small animal is named digging armadillo.
Digging armadillo is deeply adapted to a underground habit of life. The body
of animal is about 25 cm long. It has almost cylindrical shape and is covered
with strong armour. In the middle of trunk the armour of animal forms five
belts connected by elastic skin. These belts allow digging armadillo to turn
off in ball and to turn round in its own tunnels. Tail of this animal is short
and wide in the basis and has triangular outlines. Edges of corneous belts
of tail form thick strong prickles. Resting them in floor and walls of hole,
animal provides a reliable support at digging to itself.
At this animal there are short limbs. Hinder legs of digging armadillo are
rather weak and three-toed. Forepaws are wide and muscled, armed with three
huge claws (the largest claw on middle finger reaches length of 4 cm, claws
on third finger and forefinger are shorter). During the digging animal supports
on prickles on edges of tail, and rakes ground out by claws of forepaws. Friable
ground is raked back by hind legs. When the rather large amount of it is gathered,
digging armadillo turns tail under stomach, and moves back, pushing ground
to the exit of hole. On the ground surface this animal is very cautious, and
the predator practically can not creep to it imperceptibly. Digging armadillo
feels keenly vibrations of ground, and feels approaching of large animals how
they will see it.
Digging armadillo is practically blind. Its eyes are shifted in the bottom
part of head and are very small. In them there is no crystalline lens, and
the animal can distinguish light from darkness only. The head of digging armadillo
is covered with thick corneous plate in which basis thick skull bones lay.
Ears of animal are very short, protected from ground by rigid bristles. The
forepart of muzzle is protected by “visor” of corneous plate, and nostrils
are supplied with muscles permitting them to close. At digging of friable and
dry ground animal can hold the breath till 1 – 2 minutes. Due to such features
of anatomy the animal is able not only to dig tunnels but also to swim. Digging
armadillo can successfully swim across small rivers, being guided on smell
of air above water. During the swimming it rows by forepaws, and turns rear
legs in sides, and they serve for balance.
Digging armadillo eats various ground invertebrates, but the vegetative food
– roots and tubers – makes about 20 % of its diet. This animal prefers to settle
in areas with small amount of wood vegetation. Digging armadillo often builds
the common systems of holes with some local rodents, but such neighbourhood
not always passes without conflicts – digging armadilloes willingly eat newborn
rodents.
In southern regions of area this animal runs into not deep winter dormation,
does not dig new tunnels and does not leave on ground surface.
In the beginning of spring at digging armadilloes the courtship season begins.
Male is larger, than female, and behaves more aggressively. It searches in
tunnels of females ready to breeding, pairs with them and does not accept participation
in care of posterity any more. In 7 weeks after pairing in deep hole the female
gives rise to posterity – 4 cubs which always are enzygotic twins. While they
are small, the female often comes back to feed them, but since the third week
of life cubs can follow freely the female in tunnels and gradually pass to
diet of adult animals. They become sexually mature at the age of half-year.
Tiny shrew-like hedgehog (Microerinaceus minutissimus)
Order: Insectivores (Insectivora)
Family: Hedgehogs (Erinaceidae)
Habitat: New Zealand, woods in foothills.
From the beginning in New Zealand there were no small ground mammals – islands
were separated from the common body of southern supercontinent Gondwana too
early. Therefore in absence of competition one local New Zealand bat, Mystacina
tuberculata, which had adapted to partly terrestrial habit of life, replaced
them. Introducing of mammals by people had radically changed the situation
and had changed the direction of evolution of local species of animals. Among
the introduced species of mammals there were hoofed mammals, predators and
marsupials. From the group of insectivorous mammals the European hedgehog had
got to islands. It had easily accustomed in new place of inhabiting, and its
descendants had equally easily outlived humans.
The Neocenic descendant of hedgehog, tiny shrew-like hedgehog has occupied
the ecological niche of small insectivorous animals, finally having superseded
bats from it. This creature is the tiny mammal similar to shrew in size and
habits. At it needles characteristic for hedgehogs are reduced and have remained
only on head, and grow as the longitudinal strip on back. It is connected to
change of way of life: the former sluggish creature has turned in fast small
mammal scurrying among bushes and grasses; speed has replaced to it passive
protection with the help of needles. But in behaviour of shrew-like hedgehog
nevertheless there are the features inherited from an ancestor and connected
with ability to be self-protected by means of needles. Tiny shrew-like hedgehog
is rather aggressive, despite of small size. At the attack of predator even
if it is much larger than this mammal, shrew-like hedgehog is protected actively.
It bites the enemy by peaked needle-like teeth, runs into it and tries to strike
bristling needles growing on head.
Shrew-like hedgehog is colored very contrastly. Its needles are colored white,
and wool on body is black – it is warning colouring which is perceived even
by animals lack of color sight. Being protected from the enemy, the animal
utters shrill peep (almost ultrasound) which is not loved by many animals with
keen hearing. Therefore birds and mammals usually avoid to attack tiny shrew-like
hedgehog, and kill it casually in small amount. One of main enemies of this
animal is large New Zealand mouse-eating
gekko, the large lizard living in
wood litter and hunting from ambush.
Tiny shrew-like hedgehog is solitary territorial species. The individual site
is carefully marked by musk substance which is secreting from glands located
near the anal aperture. This animal is active in twilight and at night though
separate animals hunt in the afternoon, especially in shady cool woods. Males
and not sexual matured females do not arrange constant shelters and spend day
in temporary refuges. Only the pregnant female arranges a constant nest in
bush: she digs out a hole up to half meter deep, or occupies another's one.
The entrance in hole is disguised and protected by branches.
Occasionally shrew-like hedgehogs get in holes of castle
rabbits – herbivorous
animals, settling in big colonies. In holes these small mammals eat various
insects, and sometimes attack newborn rabbit cubs. But such cases are casual,
and take place only in weak colonies, where there are few adult animals.
This animal is carnivorous, and also eats only insects and other small invertebrates.
Shrew-like hedgehog has inherited from ancestor exclusive resistance to poisons;
therefore it frequently attacks even on poisonous centipeds and eats them without
harm for itself. It bites such dangerous animals in head to kill on the spot.
Frequently shrew-like hedgehog attacks lizards twice heavier, rather than itself.
In this case it carries off catch in bushes (despite of small size, it is very
strong) and stays near the catch, yet will to have eaten it completely. Rate
of metabolism at tiny warm-blooded animal is very high; therefore shrew-like
hedgehog is compelled to eat almost constantly, with small breaks on. For day
it eats the amount of food one and half times more, than it weighs.
This animal lives a little and very quickly. The female matures at the age
of about two months. After pregnancy, which lasts about 12 days, she gives
rise to 5 – 8 tiny, blind, helpless cubs. They completely develop at fortnight
age, and three-week old mammals already lead independent life. Bringing up
posterity, the female is strongly exhausted. In one month after the posterity
has abandoned the female, she restores the physical condition and is ready
to pairing again.
The close species, mountain shrew-like hedgehog (Microerinaceus montanophilus),
lives in mountains of New Zealand. It is larger (rat-sized) and more aggressive
species of mammals. Needles on its head are reduced, but on back there is a
crest of firm cross-striped needles. At danger it rears needles and stirs them
up, turning to the opponent sideways. This species also leads solitary way
of life and is a predator. It frequently attacks lizards and small mammals.
Azorean tree hedgehog (Dendrogalerix scandens)
Order: Insectivores (Insectivora)
Family: Hedgehogs (Erinaceidae)
Habitat: New Azora, lowland and mountain woods.
Usually island ecosystems differ in poverty in comparison with continental
ones. Few species can get on islands and survive in conditions of isolation.
Island ecosystems differ in vulnerability, and strongly suffer from introduction
of new species. Newly appeared species change a course of evolution of other
inhabitants of ecosystem because actually they are the new factor of environment
for native ones. But the introduced species change not less, than others, having
got in new conditions.
On New Azora Island the majority of mammalian species is descendants of species
introduced by people. The special species of insectivores, the Azorean tree
hedgehog – belongs to them. It is tree-climbing species of animals, the descendant
of European hedgehog (Erinaceus europaeus) introduced in historical epoch.
Earlier unite species had given two branches of descendants. The massive line
of ground animals is presented by large azogalerix, and small descendants of
European hedgehog turned to tree-climbing species.
Change of way of life had resulted in strongly expressed difference of new
species from ancestor in constitution and behaviour. Azorean tree hedgehog
is not able to turn in ball, and the hypodermic muscles characteristic for
ancestor became very weak at it. The body of this animal is still covered with
the pointed spikes serving for self-defense. In case of danger the animal nestles
against bark of tree and exposes towards to the enemy spikes which can rise
and fall with the help of special muscles in the basis of each spike.
In connection with tree-climbing habit of life at this animal there are relatively
larger paws, than at ground hedgehog. Azorean tree hedgehog dexterously and
quickly swarms up trees, and confidently keeps even on vertical surface of
tree trunk. Fingers at Azorean tree hedgehog are more mobile; with their help
animal can cling even to thin twigs. Besides this mammal has one more adaptation
permitting to keep on trees better. On the back side of paws at this hedgehog
especially strong and thick, slightly bent spikes grow in line. With their
help the hedgehog can cling to bark, swarming up tree trunk and branches.
Head at this animal is extended and flattened. The muzzle is extended in mobile
proboscis with which help animal searches for forage. Ears at Azorean tree
hedgehog are wide and mobile – it is necessary not only for search of prey,
but also for more effective heat exchange in conditions of hot climate. At
animal there is keen hearing due to which this hedgehog can hear noise of insect
creeping on bark in night darkness. At this animal there is good night sight,
but Azorean tree hedgehog distinguishes colors badly. The brain at this animal
is small, and behaviour differs in primitiveness.
Azorean tree hedgehog occupies an ecological niche of small prosimians in ecosystem
of this island. This is a solitary animal eating mainly small animals – insects,
tree-climbing land crabs and nestlings of various birds. Due to ability to
climb on trees this animal can attack colonies of fog
swifts settling in cavities
of trunks of large trees. Also Azorean tree hedgehog eats mushrooms and juicy
soft fruits. Among mushrooms the animal chooses the most worm-eaten ones, and
eats them along with larvae of insects settling in them. The poison accumulating
in some mushrooms does not harm to it – at Azorean tree hedgehog there is strong
immunity to poisons, inherited from ancestor.
Each individual occupies the certain territory which is marked by odorous secretions.
Out of breeding season there are no constant shelters in territory of this
little mammal. These animals meet only in period of pairing, and in another
time protect the territory from relatives. The posterity at these hedgehogs
is born two – three times per one year. In litter it may be up to 5 – 6 small,
blind and helpless cubs. Before their birth the female searches for shelter
in which it will raise cubs. Usually for this purpose it occupies hollows or
old bird nests. If necessary the female can expel medium-sized birds from liked
nests: it eats their laying or nestlings, and occupies the empty nest.
Young animals abandon nest at the age of about two months. At this time they
already see well and able to climb on branches. The litter keeps near to the
female till two next weeks, training in ways of getting of food. One-year-old
animals already can bring posterity. Life expectancy of this mammal does not
exceed 6 years.
Wormtonguer
(Vermiglossorex formicivora)
Order: Insectivors (Insectivora)
Family: Wormtonguers (Vermiglossotheriidae)

Habitat: Central and Southern Africa, savanna with small areas of wood vegetation,
wood in foothills.
When an environment after Holocene accident had stabilized, the climate of
Earth became considerably warmer and damper. At animals the huge amount
of new fodder resources had appeared, number and species variability of some
animal groups had considerably increased. In warm Neocaenic climate the area
of various social insects had considerably extended, and they became a fodder
resource for some species of animals representing new families, appeared in
Neocene.
The wormtonguer is African variant of anteater being the descendant of any
species of numerous African shrews (Soricidae). It is rather large representative
of order: cat-sized, but it seems even more largly because of features of constitution.
The head at animal is very long: it is the adaptation for feed in nests of
social insects. The muzzle of shrews is usually extended to short flexible
proboscis. This feature has received the further development at their descendant:
the skull of animal is short and expanded in front, and increased nasal cartilages
serve as support for very long mobile proboscis (making two thirds of length
of head) with nostrils and mouth on the end. Nostrils have ring contracting
muscles – it is the adaptation for protection against insect stings. Being
feeding this mammal breathes by mouth in a step to tongue movements or holds
the breath at all. Tongue is very long - it may be extended forward for length
of proboscis of animal. The basis of tongue fastens to hypoglossal bone which
has moved far back, to middle of chest. In mouth salivary glands producing
sticky saliva thickening in air are well advanced. Teeth are original – incisors,
canines and premolars are not present in both jaws, only sharp-edged molars
have remained – two pairs in each jaw. With their help animal crushes caught
insects frayed then by corneous outgrowths of stomach walls. But at young animals
at which the proboscis is not advanced yet, among milk-teeth two cutters in
each jaw develop nevertheless. Ears are short, they can fold across with the
help of special muscle. It protects them from stings of insects.
The animal spends practically all life on the ground, but if it is necessary
(for example, during flooding) it can dexterously climb on trees and even swim.
Usually the wormtonguer moves on the ground on four legs. Hinder legs are plantigrade
with straight claws and non-joined toes. On forepaws there are sharp claws,
therefore animal at walking supports on lateral side of fingers (claws at walking
are turned by edges to each other). Tail is strong, with thick basis where
the fat accumulates, assisting to go through fodder shortage. On waist of animal
there is special gland, secreting musk liquid. Colouring of wool of animal
is not striking: head of males is dark-brown colored with white tip of proboscis,
back is grey with brown cross strips, and tail is black. At females head is
grey without white spots. Claws at animals of both sexes are sandy-yellow.
Wormtonguers are active in the afternoon: from dawn till midday and some hours
before sunset. At this time insects are not so active and not so strongly bite,
or hasten to nest and are not so aggressive. Having found the nest of termites
or ants, animal rises on hinder legs, rests tail against the ground, and digs
out the house of insects by strong movements of forepaws. It licks off running
outside insects by tongue. Besides the animal can dig out of the ground larvae
of beetles and gathers single insects from plants. Usually wormtonguers live
single life, zealously protecting fodder site from encroachments of neighbours.
Borders of this territory are diligently marked by sharply smelling musk liquid.
In territory at animal there are some constant shelters, in one of which it
spends the night and will spend hottest time of day. Only in breeding season
borders of territories can be broken, and animals sometimes use this circumstance,
redistributing fodder areas. Usually one female and some males take part in
courtship ritual. Trying to impress the female, they walk on hinder legs, show
claws and loudly snort. The female couples with strongest of applicant males
though it frequently prefers most strongly smelling of them.
Two - three times per one year the female gives rise to three – four cubs.
Newborn wormtonguers are naked, blind, with closed ears. At them there are
very short proboscises, therefore they can suck milk without difficulties.
The female hides them in specially dug hole and feeds with milk about three
weeks. At three-week age young animals leave(abandon) a hole and gradually
pass to an adult diet. About one week the female teaches them to find forage,
than one week the young growth keeps near to it for protection. Later young
animals pass to independent life, searching free territory for themselves.
Young animals are not so specialized, as adults: their diet includes much more
species of insects. When the proboscis starts to grow, the wormtonguer passes
to eat extremely ants and termites. They become sexual mature at the second
year of life; life expectancy is about 10 - 12 years.
Enemies of wormtonguers are predatory birds and mammals, sometimes large snakes
and lizards attack them. The animal applies sharp claws to protect against
enemies with which it inflicts to aggressor serious lacerations. Warning about
intention to defend, animal emits odorous liquid and rubs itself’s back against
stones or tree trunks. Also, sharply exhaling air through mouth, it utters
loud sound – shrill whistle. Usually the predator has got from wormtonguer
deep wound, remembers its shout of threat and pungent smell for a long time.
Snailer
(Vermiglossorex malacovora)
Order: Insectivors (Insectivora)
Family: Wormtonguers (Vermiglossotheriidae)
Habitat: Central and Southern Africa, flatland and mountain woods.
This mammal is similar to the previous species (and it is no wonder, they are
close relatives!), but it lives mainly in damp places – in tropical woods and
thickets of bushes on river banks. The reason is, that the food of snailer
lives here - this animal eats basically snails of various species. For eating
of such food at snailer the special adaptation was developed: on the end of
its tongue horn thorns directed back have appeared. It permits to take spineless
invertebrates from their refuges and shells. The muzzle of snailer in connection
with difference in diet became much shorter than at wormtonguer, and teeth
are advanced very poorly and submitted by one pair of molars (with very thin
layer of enamel) in each jaw.
As against to extremely ground-dwelling wormtonguer, the snailer uses various
biotopes for life. More often this animal meets among trees in damp tropical
rainforest. It perfectly swarms up trees: fingers of snailer are thin and sensitive
similarly to fingers of primates. But they are armed with sharp claws, permitting
to cling, clamber and if necessary to break rotten wood. Tail of snailer is
rather flexible and partly prehensile. Colouring of this mammal is very contrast,
cryptic: the forward part of body (head, shoulders and chest) is colored black,
back one is light grey, almost white. On forepaws of animal there are white
“gloves” up to elbows and on waist and basis of tail – black round stains.
The tail of animal partly imitates colouring of some wood snakes that protects
animal from predators.
This species eats spineless invertebrates – larvae, worms and snails. Snailer
is able to break rotten tree and to take by tongue from burrows larvae of capricorn
beetles. Animal determines their presence, sniffing at apertures, bored by
larvae in wood. The most usual catch of this animal includes ground snails
making over half of its diet. Snailer takes them from shell, having cut edge
of mollusc body by claw of the thumb near the edge. After that animal pins
snail body on tongue, having pushed it in shell deeply to all coils, and by
one jerk takes mollusc meat entirely, has left shell empty.
If it is a little number of ground snails in its habitat, animals from populations
living near to rivers can dive into water, catching water snails. Having pulled
out from water such catch, snailer carefully shakes off, and then deals with
mollusc. Usually water snails, as against to ground ones, have protective corneous
operculum on their shells. Snailer cuts its contractor by claw and then eats
mollusc.
Snailers are solitary territorial animals. But as against to wormtonguers they
eat more accessible and various forage therefore they do not have sharp intraspecific
competition, and they are more tolerant concern to presence of neighbours.
As well as wormtonguer, snailer breeds some times per one year. Two times per
year female gives rise to 5 – 6 cubs. Male finds female in its territory (usually
it is the male from the fodder site located near), and some time both animals
keep and fed together. Pairing repeats some times but when female will feel
approach of pregnancy, it banishes male.
Cubs are born naked and blind, but grow quickly. At monthly age they actively
study world around, observing for mother and founding food under her supervision.
And at the age of three months they become completely independent. At the age
of one and half years female is capable to bear posterity. Life expectancy
of snailer is usually rather short – no more than 8 years. Obviously, it is
connected with fact that adult animals are practically without exception infected
with helminths, receiving them from forage – ground snails.
Ferocious condylutra (Condylutra ferox)
Order: Insectivores (Insectivora)
Family: Moles (Talpidae)
Habitat: North America, temperate and subtropical latitudes, to the north up
to Mishe-Nama Lake (southern and western coast); freshwater reservoirs of various
kinds – from lakes up to swamps.
During the evolution process low-specialized forms receive advantage in survival
first of all. Due to absence of specialization they show a wide spectrum of
variability and can occupy various ecological niches. Highly specialized species,
on the contrary, can develop only aside the amplification of specialization.
This principle is shown evidently by insectivorous mammals of mole family.
In Neocene their specialized representatives have kept characteristic shape
of digging animal, and evolution of unspecialized forms had resulted in completely
unexpected results.
In areas of temperate climate of North America one of unusual descendants of
moles of Holocene epoch lives. The structure of muzzle gives out its origin:
it is crowned up by long mobile proboscis which tip is surrounded with fleshy
outgrowths. This animal is very large descendant of star-nosed mole (Condylura
cristata) which leads semi-aquatic habit of life. The name of this animal,
“Condylutra”, combines two names: “Condylura” – star-nosed mole (an ancestor
of this species), and “Lutra” – otter (this animal partly replaces it in ecosystems).
Condylutra represents the ecological analogue of platypus and mink – it is
aquatic carnivore eating large invertebrates and small vertebrates. This is
an animal of streamline shape up to 60 cm long not including tail; individuals
from southern populations are smaller.
Head of condylutra is flat and lengthened, with low brain cavity. Eyes of this
mammal are very small, and sight is bad. But this animal searches for prey
under water not with the help of sight. Its main sensitive organ is long snout
extended to flexible mobile proboscis and covered with naked pink skin. On
tip of muzzle of its ancestor mobile sensitive shoots grew, helping to search
for food. In evolution process condylutra had further improvement of this sensitive
organ. Overgrown shoots of condylutra’s snout have electrosensitivity. With
the help of this feature animal can search in muddy water for small animals
by their electric field. External auricles at this animal are reduced, and
small ear apertures are closed by reduction of special ring muscle at diving.
Wool of condylutra is rich and velvety, colored grey. To give it the water-repellent
properties animal regularly greases hair with oily liquid secreting by two
large glands at the basis of tail. The animal finds a lot of time to look after
wool, combing it by claws of hind legs.
Condylutra leads aquatic way of life and is able to swim quickly. Diving for
prey, animal can stay under water till about 5 minutes. Short (about 10 cm
long) tail of condylutra is similar to beaver’s tail a little. It is rounded
from sides, and its back edge is dulled. Tail is covered with dense rigid skin
and thin hair; it helps to speed-up at swimming and is usually used, if it
is necessary for animal to swim quickly, not turning off (for example, escaping
from predator or banishing the congener). In this case condylutra swims with
the help of wavy movements of backbone in vertical plane. For underwater hunting
animal uses other style of swimming. Forepaws of condylutra are transformed
to narrow paddles with well advanced claws; animal uses them when it is necessary
to swim slowly and maneuverably, surveying places in which prey may hide. Diving
for food, this animal digs out bottom by claws of forepaws, overturns snags
and climbs among stalks of water plants. Also with the help of forepaws condylutra
digs at the riverbanks and lake coasts long holes with several exits. The main
exit opens right under water, and some emergency exits in case of flooding
of hole or occurrence of predator lead to shelters and to the ground surface.
Condylutra has kept predating habits of ancestor: it is exclusivaly carnivore.
Teeth of condylutra are numerous (44 ones) and pointed. This animal eats any
food of animal origin which may be found in water and near water: worms, molluscs,
crayfishes and crabs, dead or weakened fish. Condylutra can ravage clutches
and even attacks chicks of waterfowl, snatching them from under water.
It is solitary species with primitive behaviour, not forming pairs even in
courtship season. Each individual marks the territory with musky secretions
and protects it from relatives. Pairing at this species takes place in early
spring, after clearing of reservoirs from ice. In southern part of area the
repeated courtship season occurs in second half of summer. Male searches for
the female ready to pairing guiding by smell, pairs with it quickly and after
pairing the female banishes him (she is larger and stronger, rather than male).
Once a year (at the south of an area 2 times per year) the female gives rise
to 4 – 5 cubs in deep hole. The nesting chamber is always above water level.
Newborn cubs are naked and blind. They develop within 2 months, leave the female
and lead independent life. Young animals become sexually mature at the age
of 6 – 7 months: in southern parts of area young animals from the second litter
already participate in pairing and give rise to posterity the next year.
Life expectancy of condylutra does not exceed 5 years.
Long-spiked maurihystrix (Maurihystrix longispinosus)
Order: Insectivores (Insectivora)
Family: Tenrecs (Tenrecidae)
Habitat: Mauritius Island, highlands.
Representatives of ancient family of tenrecs in human epoch lived mainly at
Madagascar and Comoro Islands (from this family only African water shrews lived
in Africa). One of their representatives, rather large species, tailless tenrec
(Tenrec ecaudatus), had been acclimatized by people at Mauritius Island. After
extinction of mankind activity of introduced species had completely changed
the nature of island, and evolution at Mauritius Island has gone in other direction.
The descendant of tailless tenrec is one of species of Neocene fauna of this
island. At tenrecs there is very small brain, and this animal does not differ
in complexity of behaviour. For this reason evolution of tenrec descendant
has gone to the way of improvement of passive protection that was promoted
by fine inclinations getting from the ancestor. The body of ancestral species
was covered with wool with bristle-like spikes. This feature had received powerful
development at the Neocene descendant, and this way the maurihystrix (literally
means “Mauritian porcupine”) – the large tenrec with long spikes, similar to
a porcupine, had evolved at this island.
The body length of maurihystrix reaches 60 cm at the weight of about 5 kgs.
This is plantigrade sluggish animal with short limbs and rather large head.
Paws of animal are tenacious, and on fingers strong claws grow, with which
help animal gets various food. Head of maurihystrix is lengthened, with peaked
forward teeth and nose turned to short mobile proboscis.
Wool of maurihystrix is short and bristly, grey-colored; on stomach it is lighter.
On top of head of animal wool forms long hair “cop”. The top part of body (back,
nape, top part of neck and shoulders) of this species is covered with spikes
up to 5 cm long. Along the middle of back the crest of lengthened (up to 15
cm) spikes grows; with its help animal puts to predators dangerous wounds.
Spikes of young animals are dark, at adults they gradually brighten.
This mammal lives in mountains and other dry places, thus avoiding a competition
to Mauritian dwarf pigs, which are more moisture-loving. Maurihystrix is able
to climb on trees. Adult individuals do it very clumsily, but young maurihystrixes
climb quickly and dexterously.
This is solitary animal. Maurihystrix does not avoid relatives, but concerns
to them indifferently, not establishing hierarchy. Only at appreciable lack
of forage animals show aggression to relatives. Sight at maurihystrix is weak,
and eyes are small. For it keen hearing and sense of smell are more important,
because the animal often searches for food in high grass where visibility is
very bad. This species is omnivorous. The significant part of its diet the
vegetative food– roots and fruits of plants – occupies. Also it willingly eats
insects and small reptiles, digs grubs and worms from ground. At an opportunity
maurihystrix eats carrion.
The disturbed animal hisses and rears spikes, having turned sideways aside
the enemy. The attacked animal actively shows an opportunity of self-defense,
jumping up on four paws and having curved back like a cat.
Seasonal prevalence in breeding of maurihystrixes is not expressed. Pairing
is not anticipated by the courtsip ritual, and the female raises posterity
alone. Twice per one year it gives rise to many cubs – up to 15 ones in one
litter. Newborn maurihystrixes are completely defenceless. They are blind,
covered with thin wool without spikes. The female arranges for them a den in
rich bush, or digs a hole under roots of tree. At week-aged cubs eyes are already
opened, and wool becomes thicker. At three-week age they start to move in den
actively and try to explore vicinities. Monthly cubs can already not lag behind
the female. They abandon the nest and follow mother in wood. At this time at
them spikes start to grow. The highest death rate at maurihystrixes takes place
during the pass to independent life. The sexual maturity at young animals comes
at the age of 18 months. Life expectancy of maurihystrixes reaches 10 years.
Mole
pangolin (Talpangola laticauda)
Order: Pangolins (Pholidota)
Family: Digging pangolins (Armotalpidae)
Habitat: Southern and South-Eastern Asia, woods and light forests.
Order of pangolins is rather conservative group of mammals. From early Cenozoic,
according known fossils, they almost did not change structure and habit of
life. But in epoch of mass extinction few successful forms which have standed
“durability test” start to evolve actively, giving rise to new lifeforms.
Obviously on this wave one of pangolin species living in woods of South-Eastern
Asia, had mastered new to this group habit of life, becoming practically completely
underground animal.
Common length of the Asian mole pangolin is up to half meter including tail.
Head of animal is wedge-shaped, flattened, covered with corneous plates forming
continuous protective armour. With the help of such head animal can ram friable
wood ground, digging a hole to it. Eyes are very small, protected from above
by corneous “eyebrowes”. To protect eyes from ground at animal in addition
there are advanced dense brushy eyelashes. Auricles are not present; ear apertures
are covered by special corneous plates. Sight and hearing at mole pangolin
are bad, but sense of smell with which help the animal searches for the forage
(ground invertebrates, including social insects) is perfectly advanced. Nasal
cavities occupy significant part of skull.
Forepaws with powerful claws are the basic “digging instrument” of mole pangolin.
Claws on II and III fingers are especially advanced. Back legs are short, plantigrade,
with large feet. Almost all loading during digging and movement falls on them.
At all species of pangolins known in human epoch tails were very long. Turning
such tail around of body, pangolin could feel like in full safety from predators.
At the mole pangolin the protective role of tail is lost, as the animal spends
almost all life in holes, getting out from them only in case of emergency.
Tail of the mole pangolin is wide and short, during short travel on surface
of the ground it can protect an animal: the disturbed mole pangolin can turn
having covered belly and partly head with tail. If the enemy will not recede,
the animal makes sharp movements by forepaws, trying to wound the enemy by
sharp claws.
Body is covered with scale armour, as at all representatives of order. Between
scales it is only few wool, it is very thin. Edges of scales are covered with
corneous hairs that strengthens coupling of scales and make their abutment
more dense. Colouring of scales is rosy-brown. Because of continual friction
against ground scales of animal are “polished” and gleam slightly. Horn hairs
are formed at rubbing of corneous layer on edge of scale, and represent ends
of fibres forming every separate scale.
During digging mole pangolin serially shovels ground by claws under belly and
throws it back by legs. The main reason of special form of tail of this animal
is its use during digging holes: from time to time the animal lowers shovel-like
tail and starts to move back, raking by it away shoveled up ground to hole
entrance.
This mammal eats insects and other invertebrates licking them off by long tongue.
Basis of its diet are larvae of beetles rummaging in ground, earthworms and
snails dug in the ground. Frequently this pangolin digs burrows to termitaries,
visiting them by turns and breaking underground part of insect nests. When
from the damaged nest hundreds of “soldiers” and working individuals run out
for repair of it, animal simply licks them off by tongue, staying still.
Twice per year female gives rise to 1 – 2 cubs. As opposed to other pangolins,
cubs keep on stomach of mother - if they’ll sat on back as at ground and climbing
pangolins, female simply would dump them during movement. The female carrying
cubs, digs ground more cautiously, preferring to move in ready holes.
Cubs leave mother’s stomach at fortnight age, but about one month they are
fed with her milk. They become completely adult at the age of half-year. Life
expectancy of this animal reaches 10 – 12 years.
Lupardus (Lupardus tenuis)
Order: Carnivores (Carnivora)
Family: Canids (Canidae)
Habitat: Northern America, plains and semidesert.
During anthropogenous pressure on nature number and variety of most part of
large “noble” predators (felines, bears, canines etc.) had strongly decreased,
and some species had completely disappeared. Populations of predators were
so strongly undermined by human activity, that the most part of carnivores
species had not gone through ecological crisis at the boundary of Holocene
and Neocene, and had simply died out during several hundreds of thousand years
when on Earth unproductive ecosystems had expressed. But completely new
opportunities had been received by species which had successfully coexisted
humans. In Northern America such species was coyote (Canis latrans) which had
widely settled at the continent due to extermination of wolf and woods slash.
This species had reached prosperity in Northern America, having evolved to
some new species of predators.
One of descendants of coyote inhabits plains overgrown with grasses in the
central part of the continent, stretched like wide strip along Rocky mountains
from the north to the south. This animal is externally similar not to coyote,
and not to the representative of canine family at all, and more to someone
of felines. This similarity is reflected in the name of an animal: derivative
of words Lupus (Latin “wolf”) and “-pardus” – the ending of names of some felines.
This species is specialized in catching of swift-footed medium-sized catch
that had resulted in formation of running predator of the special appearance.
A constitution of lupardus is rather light: animal has muscled sinewy body,
thin long legs and flexible backbone. Claws at lupardus are short, but thick
and sharp: they improve coupling with ground at run. By appearance it resembles
other swift-footed predatory animals of Neocene: African pardinia and Middle-Asian
zibetonyx (both animals are representatives of civet family). Losing to other
predators in physical strength, lupardus surpasses them in speed: it is the
fastest animal of the American continent, accelerating momentum up to 90 kms
per hour at the short distance. During chasing of swift-footed catch it is
practically out of competition.
At the lupardus there is small head with short muzzle. Jaws of the animal are
rather short, but strong, with moderately long canines. Eyes are directed forward
and provide good three-dimensional sight. The nasal cavity is short, but wide,
and mucous membrane forms inside it numerous plicas. It considerably aggravates
sense of smell of predator.
During intensive muscular work the animal is exposed to serious danger of overheat.
The lupardus partly escapes from it due to fragile constitution, but it has
also extra for cooling – very wide ears similar to ears of some desert foxes.
When the lupardus chases catch, blood vessels penetrating ears, dilate, and
excessive heat dissipates. Big auricles provide to the lupardus good hearing,
that is especially important at hunting for small catch in high grass, where
the visibility is bad.
Skin of the lupardus is colored light rusty with thin close black cross strips.
This colouring resembles a little “tiger-like” colour of some breeds of domestic
dogs. Legs of lupardus are dark; strips are not expressed on them. Internal
side of wide ears is grey, but on external side of ear there is the big white
spot edged with black wool. When lupardus is turned by muzzle directly to catch,
its prey does not see the predator on the background of grass, but congeners
standing behind perfectly notice white spots on its ears.
The basic catch of luparduses is deermara – the running rodent of South-American
origin similar to small antelope. Except for them these predators hunt large
flightless ostrich turkeys. Less often luparduses attack larger animals of
American plains: donkeyhorses and young peccasons. At lack of large prey lupardus
eats small animals: rodents, lizards and snakes. Usually it avoids to hunt
among bushes or in high grass, preferring plains where grass is short because
numerous herbivores eat and tramble it. Tracking down its catch, lupardus relies
first of all on sharp sight; therefore for it the good field of view is important.
Besides the high grass complicates run, and animal simply may fall, having
hooked for stalks and having injured its legs. Having caught up prey, luparduses
put to it strong bites in sides and hips, trying to pull out a piece of meat.
With each such wound animal weakens more and more, and soon it falls, becoming
catch of predators.
Swift-footed animals have one general feature: they get tired very quickly,
and can not accelerate the maximal speed for a long time. Lupardus has the
same feature, but it easily compensates such lack by other features of behaviour.
As against cats and pardinias, and similarly to zibetonyx, it is packing hunter.
Luparduses lead the chasing of catch by organized way, and in pack always there
are animals, ready to join chasing. The pack of luparduses cuts the selected
catch from the common herd, and drives it up to exhaustion. Thus hunting animals
arrange ambushes, lining up. Herbivores cut from herd are “transferred” by
hunters from one to another. In this case the prey is every time chased by
all new animals, and by the end of chasing it is literally ready to fall down
of tiredness.
Such hunting tactics can not be organized without close interaction between
members of clan. Outside of hunting lupardus is the sociable family animal
closely communicating with congeners. Similarly to other canids, lupardus uses
for communication a tail with a white tip. By position of tail and ears animal
expresses emotion and shows the mood. The clan is ruled by main pair in which
the male is mainer than the female. Dominant male rules all males in clan and
all females submit to the main female. Youngest animals are outside of the
common hierarchy: the relation to cubs at luparduses is the most gentle and
touching. Each female twice per one year brings posterity: up to three – four
clumsy long-legged puppies. They differ from adult animals in black colouring,
and it forces even dominants to suffer all their children's pranks.
The grown up animals also outside of hierarchy, but for other reason: they
are driven off by adult animals, and young, but not developed to the full,
animals lead hard life, eating scraps. Gradually they study to hunt. It occurs
during the big hunting for large prey. In the beginning young animals are on
supporting parts, and then most capable of them quickly rise on a scale of
ranks up to the high position. And once any young strong lupardus will overthrow
the leader, proving its own hunting abilities.
Nearctic hyena-toother (Hyaenognathus macrodens)
Order Carnivores (Carnivora)
Family Canids (Canidae)
Habitat: cold and temperate areas of North America, various landscapes – from
plains up to deciduous and coniferous woods.
Canids are one of ancient groups of predators. They had made great progress
at flat landscapes being habitat of numerous herbivores. In Neocene their destiny
was changeable: having reached short-term success at plains of ice age, canids
had conceded their positions to other groups of predators better adapted for
woody districts and bushes. In such conditions canids had kept in very small
amount. Besides number of many species had been undermined by people, and they
had not gone through climatic cataclysms at the boundary of Holocene and Neocene.
But some species managed to survive.
In North America descendants of coyote (Canis latrans) had achieved the great
success in struggle for existence. Coyotes had got the big advantage from the
neighbourhood with people, having expanded the area and having got rid from
wolf, their main competitor. Possible, in first time after human disappearance
hybridization of coyote and descendants of domestic dogs took place; it had
resulted in enrichment of genetic information of new species becoming ancestors
Neocenic of North-American canids. Besides preservation of canids in Neocene
was promoted by geography of continent. Prairies formed in “wind shadow” of
Rocky Mountains, had continued the existence in Neocene. They became the house
for swift-footed herbivorous mammals and “range” for occurrence of new species
of canids. Among them two lines separated, leading radically differing habit
of life. One line of canids is represented by swift-footed packing predator
lupardus, and another one is occupied with massive animal specializing to feeding
by large catch – Nearctic hyena-toother.
Hyena-toother has occupied a convenient ecological niche of scavenger and large
prey hunter. Other representatives of American predators - large cat balam
and packing swift-footed canid lupardus catch small and swift-footed animals.
But huge peccasons, giant pigs of American prairies, are too large and strong
for them. But hyena-toothers are able to overcome such strong herbivore. This
predator lives in small packs (up to 10 – 12 individuals) consisting of dominant
pair and several subordinated pairs: they succeed to overcome large catch due
to harmonious actions. However, they easily snatch catch away at other predators
of prairies. This species effectively consumes large catch: after feast of
pack of hyena-toothers only tears of skin, jaws and leg bones remain from catch.
Other parts of prey are eaten completely.
Hyena-toother is a sizable predator (height at a shoulder is more than 1 meter)
of massive constitution: the adult animal weighs up to 300 kg. It is similar
to large and rather short-legged dog. Shoulders of animal are little bit higher
than waist, and back of animal is sloping. Colouring of animals strongly varies:
from light grey and even yellowish in southern and southwest areas up to grey
with black back and almost completely black at the north.
Paws of hyena-toother are rather short, animal can not run fast and for a long
time, accelerating maximal speed about 40 kms per hour. Toes of this animal
are rather short, and hyena-toother looks almost plantigrade. Not able to run
fast, it swims well and easily walks on marshy ground due to wide feet. Often
hyena-toothers preys swift-footed herbivores, pushing them to lakes and swamps
and chasing them on fenny ground.
Jaws of hyena-toother are adapted to feeding in carrion and crushing bones
of large animals: they are short and high, with powerful molars. Canines are
big (especially at males: it is an attribute of a sexual dimorphism) and thick,
with powerful bases. Tips of canines jut out from closed mouth of predator.
But at some external similarity to sabertooth cats it is not their analogue
by the habit of prey killing. If sabertooth cats operated by canines as by
daggers, hyena-toother operates by them differently: it puts by canines shallow
lacerations after which catch bleeds profusely.
Eyes of animal are small: sight of hyena-toother is rather weak; it badly distinguishes
colors. But the hearing and sense of smell at it are excellently advanced.
Also at hyena-toother there is high and wide nasal cavity, and olfactory epithelium
forms longitudinal plicas. Ears of animal are big, rounded and mobile.
During search of catch hyena-toothers support contact with each other with
the help of movements of tail with white hairy brush on the tip. When animals
do not hunt, they actively use sounds for communication: hyena-toothers howl,
declaring rights of clan to the territory, and also growl and whimper, expressing
the mood. Puppies can bark, but adult animals lose this ability.
The social habit of life gives big advantages to the hyena-toother in hunting
for large herbivores. Animals of different clans develop special tactics of
hunting for large animals which transfers from generation to generation. Usually
hyena-toothers choose the most perspective catch in herd of herbivores– usually
it is young or rather old animal. They rush into herd, separate the planned
victim from the common herd and drive it up to exhaustion. Hyena-toothers are
not able to run quickly; therefore they drive prey, walking behind it. They
have special places for killing catch where animals prefer to make decisive
throw and to eat catch. Usually for these purposes places fare chosen, from
which it is more convenient to reach lair and cubs.
At the period of hunting the young growth is looked after by old animals of
clan. Usually young animals start to accompany with adults approximately from
four-monthly age. Young males usually remain in parental clan at subordinated
roles with strict hierarchy, and females either pass to other clans, or form
their own one, which is leading by the male come from other clan.
Distribution of hyena-toother in North America to the south is limited by
the border of temperate climatic zone. This animal is too massive; that’s why
it badly tolerates heat. Therefore at Mexican plateau it is replaced by other
American scavenger – marsupial
hyaena (Phascohyaena tigrina), the specialized
predatory descendant of the opossum.
Anteater Fox (Neootocyon insectophagus)
Order: Carnivors (Carnivora)
Family: Canids (Canidae)
Habitat: southern and eastern Africa, savannas and woodlands
The border of Neocene and Holocene epochs was marked with the mass extinction
of various animal species and even familles. That included the extinction of
Proteles cristatus, unusual predator from southern and north-easnern Africa,
who was almost completely insectivorous. The human intervetion and new rivalring
species caused the extinction of that insectivorous hyena. But that environmental
niche hasn't stayed vacant for too long, as there were several applicants to
fill it: in the North Africa it was grumbling
ant-mungo from the family closely related to civets, while in the south
it was the representative of another carnivorous family – the bigear fox (Otocyon
megalotis) descendant – anteater
fox.
It's a middle sized animal with thick fur (size varies from 80 cm to 1 m, weight
- 6 -10 kg) and relatively long legs which is typical for most open environment
animals. Thick fur saves the animal from the ants and termites' "soldiers" bites.
Thick and shaggy fur of that animal has relatively monotone brown colour, with
just the ears' tips being a bit darker. Nose, chest and neck have extra protection
from insects, spiders and scorpions bites- unusualy dense skin. The vulnerable
nose has extra protection- a small piece of cornual "armour".Unlike
another 'anteaters' (Holocene true anteaters from Edentates Order and pangolines,
Neocene echidnas, lizards and wormtonguers) its teeth system isn't reduced-
it has plenty of sharp small teeth, resembling insectivors teeth a lot. Being
very careful, anteater fox hunts only in twilight and in the night. In the
day anteater foxes rest in selfmade holes or in the other animals dens. Very
often they share their dens with armor-headed
wart hogs for extra protection.
The sensitive ears of anteater fox are inclined a bit forward and down; keen
hearing helps it successfully find insects even in deep darkness. Its paws
aren't specialized simply for breaking termitarys, so the fox has to vary its
diet, adding another invertebtates (beetles, spiders or ants) into it. Also
its diet includes eggs and small vertebrates. One of the main alternate food
sourses are locusts, who have survived all the ecological disasters on the
border of Holocene and Noeocene. Its invasions are comparable to lemming invasions,
and as regular in Neocene as it was in Holocene. To the anteater fox, it's
just like lemming invasion to polar fox- it "forgets" about hunting
in twilight only and hunts for the locust even in the daylight.
Just like their ancestors, anteater fox are monogamous. That is explained by
the diet pecularities- they hunt for small preys so the nessesary population
density just can't be reached. The female gives birth to 3-7 cubs but not all
of them reach senior age, especially in dry lean years, when there's little
food and all savannah inhabitants, including large predators, are starving.
The maximal lifetime of anteater fox is 10-15 years.
In the northeastern Africa, where the insectivorous enviromental niche is occupied
by other animals, lives the anteater fox relative - coastal fox (Neootocyon
oraannona). Its diet includes various seafood- shellfishes, dead fish, mollusks,which
they gather during the falling tide. Unlike its southern relative, coastal
fox body is more "foxy"- it has shorter legs and smaller ears. The
lower part of coastal fox' acoustic duct is protected by skinny valve to prevent
water from coming into ears. The fur is short and hydrofuge (especially on
lower body half). Despite living at the seashore, coastal fox is not a good
swimmer. It prefers to search for its food on the littoral fringe, but sometimes
comes to feed before the falling tide ends or stays on the littoral fringe
till the rising tide starts. The fur of coastal fox is more light-colored that
the one of anteatrer fox- it's yellow with grey shades. The fingers on the
paws are mobile and broad-placed, which helps the coastal fox to run on the
sand with ease as well as hold its preys. The coastal fox is very careful animal,
who is very tolerant to its relatives. Several couples of those anmals can
feed on the littoral fringe simultaneously without disturbing each other, and
fights rarely happen during the feeding.
The breeding and home life resembles those of an anteater fox. Coastal fox
makes its dens at coastal rocks or at the bushes (higher that the highest rising
tide mark). The maximal lifespan is around 12 years.
These species are discovered
by Bhut ,the forum member.
Translated by Charles.
Ozomatli (Procyolemur ozomatli)
Order: Carnivores (Carnivora)
Family: Raccoon lemuroids (Procyolemuridae)
Habitat: North America, evergreen deciduous woods at the coast of Gulf of Mexico
and in Florida.
In Holocene representatives of primates order had gradually lost positions
occupied in prehistoric time. Only one primate – man – had achieved fantastic
power and prosperity at the expense of all ecosystem of planet. And only in
some places unique species of small monkeys could coexist with people. Other
primates became victims of hunting and human initiated change of natural environment.
And once this vicious regime had failed: the mankind had disappeared, having
fall a victim of natural laws which acted this species the same order as other
ones. Stress, illnesses and adverse heredity had undermined reproduction of
mankind as the species, and people had disappeared till some millenniums when
some waves of global epidemics had swept across the world. Taken together with
people, the part of primate species had died out: they are also subject to
human illnesses. In addition the ice age rendered large influence to the number
of primates, having reduced areas of suitable for them habitats. In Neocene
when the situation had stabilized, some ecological niches earlier belonging
to primates appeared occupied by absolutely other creatures.
One of such animals lives in woods along the coast of Gulf of Mexico. This
creature resembles a little the monkey by shape, for what it has received the
name “ozomatli” (in Aztec language it means “monkey”). Ozomatli dexterously
swarms up trees, using paws with well advanced fingers. When ozomatli yawns,
in its jaws six incisors instead of four ones characteristic for primates,
are clearly visible. It unequivocally indicates the family tree of the present
creature: ozomatli is predator very specialized to climbing habit of life,
the descendant of animal cacomistle (Bassaricus astutus), belonging to raccoons
family (Procyonidae).
Ozomatli is rather large creature: the adult male weights more than 20 kg,
the female – about 15 kg. By separate features of appearance ozomatli resembles
any monkey, but it has features, not characteristic for primates: on paws short,
but sharp claws grow instead of nails. By proportions ozomatli resembles long-legged
lemur: its head is rounded with large ears, body is lengthened, and mobile
tail approximately one and half time exceeds the body length. Paws have sharp
claws and well advanced fingers.
Rear legs are a little longer than front ones, due to what ozomatli can skip
from branch to branch. At this animal feature of structure of extremities,
more characteristic for primates, has developed: this animal does not cling
to branches by claws, and clasps them by fingers and toes. To strengthen durability
of clench, palms and feet of ozomatli are covered with wrinkled skin forming
a semblance of papillar combs characteristic for primates.
At ozomatli there is short and rather wide muzzle: animal eats mainly vegetative
food and small animals, and the part of molars had reduced. But force of bite
has appreciably increased, that permits ozomatli to chew firm seeds and to crack
nuts. Canines of this animal are also shorter in comparison with other predators.
Wool of ozomatli is short and rather thin: this creature prefers areas of warm
and damp climate, not living in woodless areas. This animal has light brown colouring
with thin cross black strips on back. Colouring of tail sharply contrasts to
the body: it is light-yellow with black-and-white hairy brush of long hair on
the tip. This hairy brush serves to the animal for communication with congeners:
ozomatlies, as a rule, are silent, and only in case of danger they utter shrilly
cry of alarm. Animals communicate with each other by means of tail movements.
Thus the hairy brush is appreciable from apart, and permits animals to not give
out themselves by cries.
Ears of ozomatli are rather wide, peaked and sticking in sides – they provide
not only excellent hearing, but also protection from overheat in warm climate.
Eyes of this animal are also big and shifted in forward part of scull. As the
muzzle of animal is short, such position of eyes provides to it three-dimensional
sight. Due to this feature ozomatli is able to jump in branches dexterously,
covering the distance up to 5 – 6 meters and more. This animal is active in day
time, as against to the nocturnal ancestor.
Ozomatli lives in deciduous woods at the southern coast of North America. It
does not avoid swamp forests of Florida and Mexican coast, but does not live
in woodless districts and light forests. This species is omnivorous and inclined
to vegetarianism. The basic food of ozomatli includes fruits, seeds and nuts
of various trees. Also this animal eats juicy stalks of orchids and other plants,
and tree leaflets. It supplements the diet with invertebrates – snails, spiders
and insects which this animal gathers in tree crones. Ozomatlies almost do not
go down on the ground, preferring to travel in crones. These creatures keep in
groups of several breeding pairs of approximately equal age, among which one
pair has leading position. Pairs at ozomatli form to all life. Partners constantly
care about each other, exchanging marks of attention: they clean each other,
constantly keep beside and exchange by silent cooing sounds with each other.
Because this animal lives in districts where change of seasons is almost not
expressed, animals having posterity may be met at any time of year. Two times
per year the female brings posterity: one large cub covered with wool. The cub
is born blind and with closed ears, and till first days of life it can only cling
to mother’s wool. It holds for wool of female, and any time she holds the cub
by one paw during jumps. At three-day age the cub starts to see and to hear,
and its grasp becomes stronger. A week later it “settles” on mother’s back and
starts to play with other cubs. Young ozomatlies keep near to parents almost
before the birth of next cub: shortly before its birth female banishes grown
up posterity, and young animals start to lead independent life. As a rule, they
leave parental clan, joining other, small group, or form groups of same aged
animals. Such groups differ in increased aggression, and can even supersede adult
animals from rendered habitable territories. Ozomatli become adults approximately
at the third year of life.
Life expectancy of this animal reaches 15 – 19 years.
Hopping carnocebus (Carnocebus saltator)
Order: Carnivores (Carnivora)
Family: South-American tail-grippers (Carnocebidae)
Habitat: tropical rainforests of South America, forest canopy.
In glacial epoch rainforests of South America had seriously suffered from climatic
changes. Because of dry and cooler climate of ice age their area had been strongly
reduced, and they had remained only along the Atlantic coast of South America,
and in lower reaches of rivers. Accordingly, the variety of local fauna and flora
also had reduced. Especially large damage was incurred by groups of specialized
wood animals among which only few species had remained. It had taken place with
earlier numerous monkeys of New World.
In Neocene the climate became favorable for growth of rainforests again, but
only few species of local monkeys had survived up to this time. In dense forest
canopy they are compelled to compete to numerous animals of other groups adapted
to tree-climbing and jumps.
One of such animals is very similar to Madagascar lemurs of Holocene epoch: it
goes to feed at night, and hides in tree-trunk hollow in day time, covering by
tail and paws large sensitive eyes. But it is not the primate, and a predator
– carnocebus, the descendant of specialized tree-climbing raccoon kinkajou (Potos
flavus). The name “carnocebus” emphasizes two features of this animal: the belonging
to carnivors order (Carnivora) and external similarity to South-American cebid
monkeys (Cebidae).
Carnocebus inhabits tropical rainforests of the north of South America. This
mammal keeps in forest canopy, practically never going down on the ground. Carnocebus
is perfectly adapted to such way of life. It has fine inheritance from the ancestor
– long tenacious tail which exceeds a little common length of body and head.
The tip of tail is covered from below with hairless cornificate skin which makes
grasp of branch stronger. When carnocebus climbs on branches, its tail, it seems,
independently grasps the next branches, providing secure if the branch appears
too thin. Paws of carnocebus are rather long because of what the animal is similar
to monkey. Rear legs are a little bit longer than front legs because of what
the back of animal is inclined forward. Due to strong rear legs carnocebus can
make long jumps in forest canopy.
Fingers and toes of this animal are supplied with short, but strong and sharp
claws. Till the climbing carnocebus simply clasps branches by fingers, but not
clings to bark, as a cat. Thumbs on hand and foot can oppose to other fingers
or toes.
Wool of carnocebus is short and velvety. General colouring of wool is brown;
on back it is darker, than on paws and stomach. Eyes are bordered by thin ring
of white wool; muzzle and area around of mouth are also white. The tip of tail
is dark, almost black.
Carnocebus has inherited the habit of life of its ancestor: this is twilight
and nocturnal animal. At it there are good hearing and very sharp sight. Ears
of carnocebus are large, rounded, covered with wool on outer side, with fluffy
fringe of long hairs on edge. Eyes of animal are big, round, directed forward:
the muzzle of carnocebus is rather short, that strengthens its similarity to
monkeys even more. Eyes of animal “shine” at night, as at cats: the retina has
a layer of cells containing guanine which additionally reflects faint light rays
to photosensitive layer of retina.
Carnocebus is omnivorous, but it obviously prefers food of animal origin. It
eats insects and small vertebrates: mammals, frogs, lizards and sleeping birds.
Carnocebus is so agile, that it can catch by jump even flying by birds and bats.
The animal eats catch, holding it in forepaws, and sitting on branch vertically.
In such moments animal additionally securing itself by tail clinging to the branch.
This is solitary animal spending day in shelter – usually in hollow of big tree.
Less often carnocebus uses abandoned bird nest as shelter. When it is placed
at top of tree, the animal arranges the original “canopy” of branches protecting
from solar beams.
Carnocebuses meet and keep for some time in pairs only in breeding season. At
this time the male some time cares at the female: cleans her fur and feeds her
from the mouth. When animals sit on branch, the male can twist by tail the tail
of the female. Animals spend day in common shelter, more often belonging to the
female. When the female feels the approach of pregnancy, she starts to behave
aggressively relatively to the male, and banishes him from the territory. Pregnancy
lasts about three months, and on the average for two years at the female it happens
three litters. It may be two cubs in litter. They are born rather advanced. Right
after birth they can cling by paws to mother’s wool, additionally twisting her
tail by their ones. They are colored darker than adult animals and have no white
marks on muzzle. When the female moves in forest canopy, cubs sit on her hips
and keep so strongly, that it can jump from tree to tree not being afraid for
their life.
Young animals keep near to the female about four months. Usually at the end of
this term the female again ready to pairing again, and cubs are banished by the
male starting to court after her. Young carnocebuses spend first months of independent
life together, and then each of them finds new territory to settle in. To this
time animals become sexual mature. Duration of their life makes about 14 years.
Taranga (Taranga partenogenetica)
Order: Carnivors (Carnivora)
Family: Mustelids (Mustelidae)
Habitat: New Zealand, woods of southern part of archipelago and mountain areas
of islands.
Before people colonization and the introducing of various species of animals
the nature of New Zealand was the most original and ancient by origin one in
the world. New Zealand is the archipelago isolated from continents since Mesozoic
era; therefore the majority of animal groups, characteristic for other areas
of the Earth, simply could not settle it. Before people occurrence mammals
of New Zealand were presented only by bats and seals. Humans had introduced
numerous mammals to islands – predators and hoofed mammals which had changed
a nature of this isolated world up to unrecognizability. Native species of
birds, reptiles and invertebrates hardly resisted to the impact of introduced
species, and the majority of them had disappeared in human epoch or during
some tens thousand years after people disappearance. Descendants of introduced
species occupied ecological niches had been empty earlier, superseded New Zealand
endemic species and after millions years had generated the balanced ecosystem.
Primary it had been no digging mammals at New Zealand, but in Neocene one of
introduced species evolved to true underground inhabitant.
One of the most successful and nocuous for native fauna introduced species
was hermin (Mustela erminea). In human epoch this mammal became the reason
of reduction of number and disappearance of numerous species of New Zealandian
birds. But time of its prosperity had approached to the end in glacial epoch,
when the climate had changed. Ecosystems of the past degraded, and inhabitants
of islands should search for new sources of forage to survive. One of descendants
of hermin had turned to digging animal, having replaced moles absent at these
islands. Taranga – the new species of digging mammals – had appeared so in
New Zealand. In myths of New Zealand Taranga is the female, in the afternoon
leaving the land for the underground world. This animal, as against the mythical
prototype, had gone under the ground for ever, becoming the species deeply
specialized to underground habit of life.
Body length of taranga is about 20 cm, from which about a quarter falls to
a tail. The body of this small mammal is covered with rich velvety wool of
grey color. Populations from mountain areas differ in longer and denser fur.
Head of this mammal is very short and flat; on nose bridge the wide corneous
scute is advanced. Taranga digs holes with the help of forepaws. They are similar
to paws of the mole: short, wide, strong, with palms turned back and long thick
claws. The body of taranga is lengthened, hinder legs are rather weak – animal
pushes by them from walls of hole during the movement.
The skeleton of this mammal is additionally strengthened: it is important for
underground habit of life when the animal is threatened constantly with danger
to appear filled up with stones. Cranial bones of taranga are very thick; ribs
are expanded and strong. Vertebrae are strengthened by numerous additional
shoots and jags forming additional mechanical coupling. By structure of backbone
this animal is convergently similar to girder-backed shrew (Scutisorex somereni)
from Uganda lived in Holocene epoch.
Sense organs at taranga are advanced differently, rather than at hermin. Eyes
of animal are very small; sight sence is substantially degenerated: the animal
badly distinguishes contours of objects and does not distinguish color at all.
Sight is useless at life under the ground, and by importance for animal it
had been completely replaced with well advanced sense of smell.
Sharp thin teeth specify that taranga is active predator. This animal eats
rodents, worms and reptiles creeping in its holes.
Taranga does not like stony soils and lives in soft wood ground rich in humus.
Separate populations of taranga exist in mountain areas of New Zealand, but
they are settled in places where there are woods with thick layer of ground.
Also taranga does not live in tropical woods of the north of New Zealand, where
the layer of ground is rather thin.
This is the solitary animal. Each individual digs system of holes and marks
borders of territory with musk secretions. If two animals meet in the common
system of holes, there is a fight between them, frequently ending by the death
of one of animals. For these animals the cannibalism is characteristic, especially
at the lack of food.
Only the female ready to pairing can admit the male to the territory, but right
after pairings she banishes him. At this species of mammals the number of unique
features of breeding is developed. The hermin – the ancestor of taranga – strongly
depended on number of rodents, and at it the special adaptation permitted to
restore quickly the number of species had developed. The male of this species
had coupled to newborn females, and they grew, being pregnant. At taranga this
adaptation had undergone change in connection with warming in early Neocene
and rather constant conditions of inhabiting. This animal is able to parthenogenesis:
young females grow, and at them the new parthenogenetic generation of cubs
already develops. All newborn individuals of parthenogenetic generation are
females. If the female ready to pairing has not met the male, at her the occurrence
of parthenogenetic pack is also possible. In ovocites there is the doubling
of chromosomal complement which is not accompanying with cell division, and
from them normal cubs develop. In mountain valleys there are the populations
of taranga including only females – possible, this is the parthenogenetic posterity
of the single foundress individual. Taranga breeds two times per year; in litter
it may be 4 – 5 cubs.
This animal grows quickly, reaching the maturity at the age of half-year. But
the life expectancy of taranga exceeds five years seldom.
Taranga is not unique species of underground predators on Neocenic Earth. At
the territory of Europe the blindweasel lives – it is another species of underground
mustelid, completely lost sight sence, but not adapted to burrow digging.
Gliding weasel (Ventomustela zewanna)
Order: Carnivores (Carnivora)
Family: Mustelids (Mustelidae)
Habitat: Balkan, woods to the north of the Alps; forest canopy.
In Neocene subtropical woods had returned to Europe. But trees forming them
were not natives from the south. Mediterranean Sea, the shelter of thermophilic
flora, had turned to heated salt desert, and the rich southern flora was lost,
having replaced by drought-resistant plants of desert. Trees of European forests
are descendants of species of flora of temperate latitudes: broad-leaved oaks
and maples, relic plane trees, nut trees, beeches and chestnut trees. They
had managed to survive in mountain valleys during the ice age, and after restoration
of conditions favorable for life, had formed rich shady forests, stretched
by strip from the Central Europe (to the north of the Alpes) through Tatra
Mountains and Carpathian Mountains up to coast of Fourseas.
On the ground under forest canopy a few kinds of animals live, and the majority
of inhabitants of European deciduous forests live in forest canopy. As against
tropical rainforests, in European subtropical forests there are only few species
of lianes, and crones of separate trees are not connected by them together.
Therefore in canopy of European forests only ones live, able to fly well or
… to make long gliding jumps. Abundance of prey had involved to forest canopy
the predator perfectly adapted to movement in such inhabitancy. It is the representative
of mustelid family, the gliding weasel.
The European gliding weasel has kept small size, characteristic for many mustelid
species. The length of body of this animal is about 30 cm, and about 20 cm
falls to fluffy tail. Between paws of gliding weasel skin membrane is advanced,
permitting this little mammal to make long gliding jumps from tree to tree.
Head of animal is small and rounded, with short ears. Eyes are shifted to forward
part of skull and provide good three-dimensional vision – it is vital for animal
living in a forest canopy to estimate distance before a jump more precisely.
The short wool of gliding weasel is colored brown, and on tail there are some
black rings. The bottom part of body of gliding weasel is snow-white. To winter
at individuals in northern part of area wool turns longer and a little lighter.
This animal is active and successful predator (it has received the specific
name in honour of goddess of hunting of western Slavs). At gliding weasel there
are large canines, and mouth can open very widely. Gliding weasel eats various
forest rodents, bats and birds. This predator kills prey by bite in head. Gliding
weasel often ravages bird nests, showing miracles of acrobatics and dexterously
landing even on very thin twigs where the nest is built. This animal can climb
in hollows to bee nests in searches of honey, and eats other insects and large
spiders. Gliding weasel spends not whole life in tree crones. It often goes
down on the ground in searches of forage. Being a predator by its nature, it
does not disdain ripe berries, and in autumn cracks large oily seeds of beeches
and chestnuts which grow in forests of Southern Europe much. On the ground
this animal eats worms and insects. This mammal is able to climb on trees as
dexterously, as squirrel, despite of presence of gliding membrane.
Gliding weasel is a solitary animal. Each individual has site in forest and
marks it with odorous musky secretions. The territory of animal includes not
only a part of crones of trees, but also trunks and the ground under trees.
Gliding weasel pair two times per year. First time courtship games occur at
the end of winter when night frosts stop. Males start to come in territory
of females and to leave their own odorous marks near female’s ones. If the
female is ready to pairing, her musky secretions get a special smell well distinguishing
by males. When the male feels it, he tries to reach the female as soon as possible.
Sometimes at once some males follow one female. They constantly compete with
each other, and sometimes can bite each other, putting deep painful wounds.
Sometimes competing males seize each other’s gliding membrane, and one of them
can pay for failure in courtship season by loss of ability to make gliding
jumps. Eventually, near the female only one male stays, which pairs with her.
After the pairing the female banishes the male. Pregnancy lasts till about
two months, and in second half of spring the female gives rise to 4 – 6 cubs.
They are blind and helpless, and female looks after them within 2 months before
they become independent. She hides cubs in deep hollow of tree at the big height.
Young gliding weasels differ from adults in darker colouring of wool and absence
of strips on tail.
Second time for a season the female is ready to pairing approximately in one
month after cubs had ceased to suck milk. The first days of pregnancy the female
concerns to grown up cubs tolerantly, but later starts to express aggression
to them more often, and compels them to live independently. Second time it
brings posterity at the end of summer. In the second litter it is less number
of cubs – only two or three ones. But they are larger, than ones in first litter,
and become quite independent predators to the beginning of winter.
The main enemy of these animals is large gospodar
woodpecker which pecks out
tree-trunk hollows and can eat newborn cubs of gliding weasel.
The idea about existence of this animal was proposed by Momus, the forum member.
Antipod’s unotter (Xenolutra antipodorum)
Order: Carnivores (Carnivora)
Family: Mustelids (Mustelidae)
Habitat: New Zealand, sea coast and lower stream of rivers.
Freshwater ecosystems of New Zealand differed in poverty and high degree of
endemism in comparison with continental ones. It is connected with very early
isolation of archipelago, which had taken place before occurrence of the most
typical families of freshwater fishes and other water vertebrates. The situation
had sharply changed in human epoch when various species of freshwater fishes
and predatory mammals had been introduced to islands. After disappearance of
mankind their descendants had changed freshwater ecosystems up to unrecognizability.
One of such species lives in lower reaches of rivers of New Zealand; it is
a predatory aquatic mammal, crowning with itself the food pyramid of ecosystem.
By shape this animal is very similar to otter, but is connected to it with
only remote relationship by origin: it is the descendant of the domesticated
American mink (Mustela vison) introduced to islands in human epoch. This animal
is named antipod’s unotter. It is much larger, than the ancestor: up to one
and half meters long including wide flattened tail.
At antipod’s unotter there are rather short paws with long fingers connected
by elastic membrane. The antipod’s unotter is very vulnerable on land, but
in water only few animals can make a competition to it. For slow and maneuverable
swimming animal uses paws, but at hunting in open water unotter presses paws
to body and swims with the help of vertical bends of backbone and movements
of tail. Tail of antipod’s unotter is wide and flat, similar to beaver’s one.
It is covered with dense skin and shorter fur, than the body.
Antipod’s unotter has flattened head with short strong jaws which easily crack
skull of fish up to 40 – 50 cm long, or the armour of large crab. Ears of animal
are short and rounded. On muzzle of unotter long rigid whiskers – vibrisses
– grow; with their help animal searches for prey under water.
The body of unotter is covered with dense rich fur of grey color with white
spot on throat which passes to longitudinal strip on stomach. Fur of antipod’s
unotter is rich and dense. It reliably keeps air, and consequently animal is
not cold even in cool water. Unotter carefully looks after fur, and combs it
with two wide claws of forepaws which bottom surface forms “combs”. The animal
also greases fur with secretions of former musky glands, located near the anal
aperture. In this connection they had strongly changed: having lost smell component
(the strong smell is characteristic for many mustelids), they became plentiful
and oily. Cubs at which glands poorly function yet, use for care of fur secretions
of the female.
Antipod’s unotter is active piscivorous predator. It usually hunts in coastal
zone and in thickets of brown algae. These animals especially willingly catch
pelagic fishes which shoals swim up close to the coast. The group of animals
attacks fishes, and drive their shoal into algae thickets where fishes lose
orientation: in such situation it is easier to catch them. Besides of fish,
antipod’s unotter eats crustaceans and small snails. After storm these animals
gather on coast dead sea animals cast ashore.
By the way of life the antipod’s unotter is social species. It settles at sea
coast or in mouth of the river in big groups numbering up to ten breeding pairs
of adult animals. Besides of them in groups there are some single adult individuals,
and also numerous youngsters – the posterity, yet not completely passed to
independent life. The colony lodges in places where there is an opportunity
to arrange shelter. Animals dig in the ground some common holes, or lodge in
cave well protected from storm waves.
The pair at antipod’s unotters is formed to all further life. Partners differ
in attachment to each other – they have a rest and hunt together. Male is larger,
than female and female regularly renders to him signs of attention and submission
– she clears and greases his fur and licks his muzzle. Once a year, in early
spring the female gives rise to 3 – 5 cubs. They are born blind and deaf, but
covered with wool. The female gives rise to posterity in shelter, on litter
of grass and seaweed. First some days she does not admit to posterity even
her male. Week-aged cubs start to hear, little bit later eyes open at them.
At the age of 17 days cubs start to creep out from den and to explore the world
around. The female feeds them with milk approximately up to bi-monthly age,
but already fortnight cubs start to try food of adult animals: male brings
in the den fish, and the female allows cubs to eat a little. At the age of
two months young animals already leave a den and start to study to swim and
to get food independently.
As a rule, males abandon the group at the age of half-year. They lead barchelor
life, and can found their own colony if they find a place suitable to life.
If the parental colony is overpopulated, adult individuals can show aggression
to young growth. Then even young females leave a colony; later they can join
a colony of bachelor males and form breeding pairs with them. At the age of
two years young animals become able to breeding.
Berl (Quasiursus asiaticus)
Order: Carnivores (Carnivora)
Family: Mustelids (Mustelidae)
Habitat: Eurasia, North America, temperate and cold areas, from the north of
Europe up to Greenland.
Large species of carnivorous mammals are the most vulnerable species in an
ecosystem. At strong (though and convertible) fluctuations of number of prey
number of predators can fall down to critical level, and the species appears
at the edge of extinction. Development of extensive territories of land by
people, and the destruction of habitats followed it had strongly reduced number
of large predators, and they had not gone through ecological crisis at the
boundary of Holocene and Neocene.
After extinction of mankind efficiency of ecosystems began to restore. Numerous
descendants of small species, mainly rodents and lagomorphs had occupied the
place of large herbivores exterminated by people and had become extinct because
of natural reasons. After herbivores, some predators also had strongly increased
in size. Among large predators the serious application for domination in fauna
had been made by representatives of mustelid family. In temperate and partly
in subtropical zone of Northern hemisphere their representative, berl, is found.
It is a huge carnivorous species of mustelids, the analogue of fossil marten
Perunium, and the descendant of ferret (Mustela putorius). Its ancestors, probably,
took in ecosystems a place similar to wolverine (Gulo gulo), and any of their
species had managed to enter large-sized class. This attempt appeared extremely
successful.
Berl is massive and sluggish species of mustelids, in size compared to bear
(hence the name: “berl” is the ancient Slavic name of bear). The height of
adult animal at a shoulder exceeds one meter, length of body without tail is
up to three meters; weight is over 400 kgs. The adult animal, as against bears,
hardly swarms up trees (cubs, on the contrary, do it very dexterously). By
constitution berl resembles polar bear (Ursus maritimus) of human epoch – it
has rather small head on strong neck, large trunk and muscled paws. As against
bears, berl has long fluffy tail. It is solitary and aggressive species of
mammals, and by position of tail each individual expresses the mood, not entering
close contact to relatives. Fur of berl is rich, especially in winter. Summer
colouring of wool of animal varies from dark grey up to beige, straw-colored
and even black. Usually lighter-colored individuals live at the south of area,
and darker ones – in forests of temperate zone. These animals are able to change
color of wool to winter. Individuals from cold areas become snow-white in winter,
and in more southern areas winter fur of animal turns grayish. Individuals
from southern borders of area are almost unable to change color of winter wool
– it becomes thicker and only a little lighter. In winter berl keeps activity
even in ringing frosts.
Holocene bears were omnivorous and even mainly herbivorous ones (except for
polar bear). Berl is an exclusive predator. Using huge physical strength, it
attacks large mammals, making the basis of its diet: in Eurasia obda, aurochid,
nozdrokh, shurga, in North America giant
wood porcupine, in Greenland snow
porcupine and skewhorn. Berl can not chase its prey, therefore it uses another
tactics – it pursues herd of herbivores during the long time while the weakest
animal will keep abreast. When prey appears far enough from relatives, berl
rushes on it and kills by impact of paw – it simply brings down animal on the
ground and breaks its neck. Hunting takes a lot of time and forces, therefore
berl aspires to use food resources maximum full. This predator stays near killed
prey for a long time: in summer while meat will start to spoil, and in cold
season while on carcass there is at least something edible. Also berl willingly
eats carrion, or takes by force prey of smaller predators.
This predator is solitary animal; each individual has extensive fodder territory.
Berl marks borders of territory by unpleasantly smelling liquid which is secreting
from anal glands. The animal leaves marks on stones and trunks of trees. In
courtship season (in the beginning of autumn) males wander on territories of
females, and couple with them. Only at this time females become more tolerant
to presence of adult relatives in their territory.
At the end of spring when it becomes warm enough, the female arranges in forest
carefully disguised den in which brings posterity: two – three blind and helpless
cubs. The posterity develops rather slowly: only at bi-monthly age cubs start
to leave den and to explore the world around. They spend a lot of time in games
– swarm up trees, combat and simulate hunting, “attacking” on trunks of trees
fallen on the ground. In autumn the posterity finally abandons shelter, and
studies to hunt. Young animals keep in common with the female during all next
year and all this time demand attention from the part of mother. Therefore
the female of berl brings posterity only once in two years.
Berl is widely settled in Eurasia and at the north of North America. It lives
mainly in areas of coniferous and deciduous forests, forming a number of subspecies.
At the territory of Siberia and Far East very large nominative subspecies great,
or Siberian berl (Quasiursus asiaticus asiaticus) is found. At this subspecies
there is the darkest summer colouring of fur, and winter fur is greyish-white.
At the north of Europe it is replaced by smaller European subspecies, mechka*
(Quasiursus asiaticus occidentalis) which has the smaller size and reddish
or yellowish colouring of summer fur. The north of Eurasia and North America
is occupied by indistinctly separated from these forms subspecies oshkooy**
berl (Quasiursus asiaticus leucos) with snow-white winter and dark grey summer
fur. This animal lives in woods of Far North, often hunts and gathers carrion
at the coast of Arctic Ocean, and migrates on ice to northern islands in winter.
At the south of area this kind forms hybrid forms with European and Asian subspecies.
At the territory of North America the Asian berl is replaced by two close species.
Mountain berl, or ueb (Quasiursus montanophilus) lives in woods of Rocky Mountains
ridges, coming along high-mountainous areas far to the south (up to territory
of Mexico). This species dark grey summer fur, often with white spots on lower
jaw and breast; at some individuals there is white stomach. This species is
only a little smaller than Siberian berl, especially the representatives of
northern populations distinguished by shaggy rich fur. Winter fur at this species
is bluish-grey. Ueb differs from Asian berl in larger head and more flat teeth
– it is appreciably less selective in food. In summer this animal often eats
vegetative forage – mushrooms and berries.
Eastern berl (Quasiursus appalachensis) is much smaller than its western relative,
and is approximately equal in the size to European mechka. It is colored yellowish-brown
color and keeps such color of wool the year round. This species differs in
lightest constitution among the relatives, and swarms up trees much better.
It occupies evergreen and semi-deciduous forests of the eastern part of North
America.
In Eastern Asia there is one more species of this genus - tiny berl
(Quasiursus minimus). This animal is the analogue of wolverine, medium-sized carnivorous
species (its growth at a shoulder is no more than half meter). It inhabits
forests of various types, woodlands and mountain forests, and also boggy woods
in river valleys. This animal eats various mammals, large birds and fish. It
is able to climb on trees, to swim and to dive. The skin of tiny berl has pale-yellow
color, and paws are dark brown; on muzzle there is dark “mask”.
Red berl (Quasiursus rufus) living in Japan is close to this species. It is
larger, than tiny berl (growth at a shoulder is about 60 cm), more thickset
and strong combined. At this animal there is rich rusty red fur and wide feet
– it feels well in high mountains and easily moves on snow, not failing down.
Red berl lives in northern part and mountain areas of Japan Islands, preferring
woody districts and thickets of bamboo. It also is a predator, and prefers
to hunt mammals.
* “Mechka” is also Slavic name of bear;
** “Oshkooy” is the name of polar bear at folks of the North of Eurasia.
Neosurilio (Neosurilio zebropygus)
Order: Carnivores (Carnivora)
Family: Mustelids (Mustelidae)
Habitat: forests of the east of North America and along the coast of Gulf of
Mexico.
In epoch of biological crisis at the boundary of Holocene and Neocene the species
had not having strict specialization received advantage in survival. The part
of these species belonged to the group of synanthropic ones, and had adapted
to live near people, having received from it additional benefits – an abundance
of forage, refuge and rather quiet existence. But after human disappearance
all species had received equal opportunities for further evolution.
Among mustelid family among survived ones there were North American skunks
– the omnivores having an effective way of protection against enemies. One
of descendants of spotty skunk (Spilogale gracialis), neosurilio (“surilio”
is the name of one species of skunks in human epoch) lives in Neocene epoch
in forests of North America. It is the ecological analogue of badger, the sluggish
plantigrade animal having massive constitution. Length of body of this animal
is about 70 – 80 cm, and length of tail is about 40 cm.
Neosurilio is noticeable from apart due to contrast warning colouring. The
animal as if consists of two halves sharply differing by colouring. On back
part of body, hips and waist there are alternating black and white strips (hence
the scientific specific name meaning “having back of zebra”). Tip of tail is
also white. Forward part of body of neosurilio is brown with white throat and
stomach. Around of eyes there are “glasses” of white wool, and on top of head
there is black “cap”.
Neosurilio has kept way of protection characteristic for skunk: it defends
against enemies splashing smelly liquid from strongly advanced anal glands.
In special sacs the oily liquid containing ethylmercaptane accumulates in amount
enough to make some “shots” in succession to distance up to 3 meters. Before
application of the weapon animal shows to aggressor the black-and-white back,
having turned it to aggressor. At aggressively behaving neosurilio wool on
back part of body rises on end and fluffs, increasing a little the visual sizes
of this part of animal. If this warning had not worked, jet of smelly liquid
flies to predator.
Neosurilio consumes various kinds of food, and due to the omnivorousity it
does not stay hungry in any district. On forepaws of animal thick strong claws
grow, with which help neosurilio easily digs out holes of small animals and
gets roots and tubers of plants. More often neosurilio eats small animal –
invertebrates, amphibians; also it ravages nests and eats eggs and nestlings
of small birds. Up to half of diet of this animal the vegetative food makes
– tubers, roots of plants and berries. Neosurilio eats food “aristocratically”
– it sits on hind legs and brings it by front paws to the mouth. At the end
of summer and in autumn neosurilio almost completely passes to vegetative food,
and during winter and spring food shortage does not squeamish about carrion.
Using the chemical weapon, neosurilio behaves very impudently, and can drive
away from prey even large predators.
Each individual builds in the territory some temporary shelters and spends
the night in them during wandering on the territory. Also neosurilio does not
overlook to mark border of territory, splashing some drops of combat liquid
on roots of large trees. The female ready to birth of posterity becomes homebody.
Right before birth of posterity she stays best arranged shelter, and starts
to expand it even more. She digs a hole up to 5 meters deep with several emergency
exits. Pregnancy at neosurilio lasts about three months, and proceeds from
the end of winter up to middle of spring. The female gives rise to 2 – 3 large
cubs. They are born blind and deaf, but already at birth are covered with wool.
The female cares of them for a long time, feeding them by milk up to 4-month's
age. But to this time cubs already develop enough to leave a hole and to follow
the female. At this time at them also odorous glands develop and they become
able to splash smelly liquid. Young animals spend the first winter with the
female, learning the search of food, and leave her shortly before courtship
season when she starts to behave aggressively to them.
Ursine
civet (Ursivetta aromatica)
Order: Carnivores (Carnivora)
Family: Civets (Viverridae)
Habitat: mountain woods of Southern Asia.
Mass extinction of live creatures at the border of Holocene and Neocene practically
had not given chances of survival to animals weighting more than 20 kgs. Predators
dependent on abundance of large prey – cats, dogs, hyaenas and bears have especially
suffered from it. But small predators like raccoons, weasels and civets have
easily survived and have “privately divided” the planet: weasels and their
descendants became main predators of temperate and cold areas of Earth,
and civets were spreaded in tropical zone of Old World. Only in Africa and
partly in America relic representatives of cats, descendants of small cat species
compete with them. Holocene representatives of civet family by the constitution
are analogues of primitive predators of early Cenozoic, therefore it is possible
to assume, that they have the big evolutionary future. Among them there are
ecological analogues of omnivorous bears and carnivorous cats, therefore it
is possible to assume, that in the future they will occupy place of these animals.
Ursine civet is the huge representative of civet family. This animal is similar
to bear with long tail (prehistoric analogue of ursine civet is the huge fossil
marten Perunium dwelt in Eurasia). Constitution of this animal is, however,
more graceful, than at bear: neck is longer and more mobile, and muzzle is
more extended. Adult animal weighs up to 250 - 300 kg (males are larger). Legs
are almost plantigrade, with long non-retracting claws. Fur of animal is rich
and rather long (the animal frequently meets in mountains, sometimes in summer
rising up to zone of Alpine vegetation), chestnut-brown with mesh white pattern
on neck and back. On tail there are some cross rings of white wool, the tail
tip is black. With the help of tail position ursine civet expresses its mood
at the meeting with neighbours. In the basis of tail there are repugnatorial
glands which emit pleasant musk smell for territory marking.
This animal is solitary one living in territory numbering some square kilometers
in area and strictly protected from neighbours. Only female and cubs can live
any time in group.
The ursine civet is omnivorous species with deviation to predating. Teeth at
it are wide, knobby and dulled – vegetative food makes about 40 % of diet.
Among plants ursine civet prefers fruits of different palms rich in oil. Also
it digs out by claws roots of huge grasses of family Apiaceae growing on mountain
slopes, and tubers of various wood grasses. On river banks ursine civet eats
roots of reed mace and other marsh plants. The food of animal origin includes
larvae of various insects, frogs and lizards, eggs and nestlings of ground
birds. If there is an opportunity, ursine civet willingly eats carrion and
attacks cubs of ground animals. Due to claws this animal can climb trees, ravaging
bird nests.
Ursine civet is mainly day time animal. It spends night in shelter which it
is usually a little number in its territory. From time to time animal moves
to other shelter, giving to ants and the other insects the right to eat staying
in lair litter larvae of fleas and ticks. For disposal of parasites animal
willingly bathes in river and wallows on odorous grasses, especially on greenery
of plants of Apiaceae family (having eaten its roots).
Borders of territories can be broken only in season of pairing. Usually on
border of possession of several animals there is special remarkable place like
alone growing tree or large stone. Animals make here original “message board”,
marking it with odorous secretions. When the male notices on smell, that the
female is ready to pairing, he puts the mark atop of mark of this female, and
waits nearby from “message board” driving away other males. Also he puts marks
in territory of chosen female: peels bark from tree and marks this place with
odorous secretions, declaring about its presence. When the female comes to
“message board” and sniffs at it, she becomes acquainted “in absentia” with
the male, and if the male is pleasant to her, some time it allows him to remain
nearby for pairing.
When the female feels approach of pregnancy, it banishes out the male, sometimes
should win anew his own territory if on it in his absence the neighbour had
lodged. Pregnancy lasts about five months. Shortly before cub birth the female
builds lair – she digs out hole under tree roots or finds ready cave in mountains.
She drags moss and branches for litter, and carefully hides traces of her presence
near to chosen refuge.
In pack there are usually two cubs though sometimes there are triplets, and
young females bear only one cub. Young growth is born blind and with closed
ears, covered by short wool. Colouring of newborn cubs is spotty: chestnut
stains on red or straw-coloured background (characteristic colouring of Asian
civet (Viverricula), an ancestor of the ursine civet, was spotty), later cubs
darken. At ten-day age at them eyes and ears open, and month-aged cubs already
can play near lair entrance. At bi-monthly age cubs leave shelter, and female
teaches them in ways of food getting. Young growth stay with mother up to eight-monthly
age then starts to live independently. Sexual maturity comes at them at three-year
age, and life expectancy can reach up to 30 - 35 years.
Pardinia
(Pseudacinonyx gracilis)
Order: Carnivores (Carnivora)
Family: Civets (Viverridae)
Habitat: savannas and light forests of Southern Africa.
Mass extinction of large herbivore animals has caused disappearance of large
predators eating them. After stabilization of situation and restoration of
biological balance as the reply to evolution of herbivores large predators
have started to appear – they were descendants of rather small forms which
have survived during mass extinction.
Pardinia is one of such predatory animals, the descendant of genette (Genetta
genetta), widely spreaded in Holocene at the territory of Northern Africa and
Southern Europe. Its close relative is the huge deadlynetta (Necrogenetta
deima), living in North-African savannas. But if deadlynetta is the specialized
sabertooth hunter for large prey, pardinia is graceful analogue of the cheetah
chasing swift-footed herbivores. By constitution pardinia resembles fleet dog
with long tail and rather short muzzle. Height of animal at a shoulder is about
80 cm, body length is up to 120 cm, tail lenght is about meter; back legs are
little bit longer than front ones. Body of pardinia is covered with short wool,
on edges of ears there is fringe of long soft hair. Wool is straw-coloured
with scattered brown stains on back merging to longitudinal strips. On throat
of males there is big white spot bordered by dark-brown strip. Head of pardinia
is similar to cat’s one, but is more extended. Canines are well advanced, their
ends are visible from closed mouth of animal; bases of canines are very wide.
Chewing muscles are well advanced: animal kills catch by bite in backbone or
in the basis of skull.
Similarly to cheetah pardinia chases catch by fast throw. But as against cheetahs
it also can chase lonely catch walking for a long time, yet will drive it up
to exhaustion. Chasing prey, pardinia tries to fall catch down on the ground
by impact of paws against side or shoulder, and then presses it by own weight
against ground and bites in backbone.
Pardinia is solitary predator. Each female has territory marking it with secret
of musk glands. Male, as a rule, has no constant territory: he wanders on territories
of several females, remaining for the pairing period at one of them. Pregnancy
lasts 5 months. Shortly before cub birth female builds shelter in rich bushes
or among stones where it gives rise to posterity: 4 - 6 cubs. Cubs are born
blind, covered by dark one-colour wool. Later their colouring replaces to motley
one, characteristic for adults. Cubs develop rather slowly: they start to leave
shelter at monthly age, and study to hunt approximately from three-monthly
age. About half-year they hunt together with mother, leaving her shortly before
occurrence of new pack. Sexual maturity at young animals comes at the second
year of life.
Pardinia hunts mainly quickly running herbivore animals - hoofed mammals, kept
in Southern Africa as relicts of Holocene fauna. Hunting usually occurs on
dawn though the female having cubs can hunt even in hottest time of day. Pardinia
creeps to herd hiding in bushes and grasses, stopping at slightest sign of
alarm at herbivores. Having chosen prey animal tries to catch it by one throw
– pardinia is not able to run fast at long distance. Usually only every 6-th
or 7-th attack ends successfully.
If number of large catch is not enough, pardinia willingly foods with small
vertebrates – frogs, lizards and also birds. Dexterity of the pardinia is great:
it can drop flying up birds in jump by long paws.
Organism of pardinia as against to organism of deadlynetta related to it works
literally “for deterioration” – during run plenty of forces is spent. Therefore
life expectancy of this animal is not so great: in total up to 12 – 14 years.
Long-bodied pterogenettula (Pterogenettula prolonga)
Order Carnivores (Carnivora)
Family Civets (Viverridae)
Habitat: Equatorial Africa, the top level of rainforest.
Development of various anatomical adaptations at live beings depends on features
of their environment. And the generality of inhabitancy results in formation
of similar adaptations at unrelated species. It has taken place in woods of
Equatorial Africa where on top-sized trees many species of mammals able to
make long jumps were developed. If the forest canopy is dense, in it tree-climbing
animals receive advantages. But in friable multileveled canopy experts in long
jumps dominate. And on top of food pyramide there is also living “glider pilot”,
but it is a true predator.
Predator of the top level of African woods is the long-bodied pterogenettula,
small animal with extended body. It had descended from one of species of wood
civets managed to go through the time of active hunting in last equatorial
woods of Africa.
Body length of this predator is about 30 cm at all, and tail lenght is about
20 cm. Externally this animal is similar to any small weasel: at the pterogenettula
there are rather short legs and the extended head. The animal has “marble”
spotty colouring of large ring-formed spots on light background. It makes this
predator imperceptible on the background of bark spotted with shadows of leaves.
On head there are dark strips between eyes and nose, needing for recognition
of neighbours.
Extremities are rather short, supplied with tenacious claws. But the most remarkable
feature of appearance of these small mammals is the presence of small flying
membrane between front and hind legs. For improvement of its “flying” properties
on edge of membrane the fringe of black fluffy fur (as on sides at Holocene
monkey Colobus) is stretched. The long tail of animal serves as rudder. Skilfully
using force of wind, the pterogenettula can make jumps up to 20 meters long,
losing in height only 8 - 10 meters. Operating flight, the pterogenettula can
even catch small birds, preying them in sudden jump from an ambush. If attack
was unsuccessful, the animal simply continues a jump, and sits on a tree. Flying
up to the final point of jump, the animal weakens flying membrane, and seizes
by claws of all four paws branch or bark of tree. Also this predator is able
to swarm up trees dexterously, being not inferior to any squirrel in speed.
The pterogenettula frequently hunts also a way traditional for predators, chasing
catch among branches.
Favourite catch of the pterogenettula includes birds, but also it frequently
hunts lizards, squirrels and squirrel
guenons - dwarf monkeys of forest canopy.
This animal is strict solitary predator. Each individual chooses to itself
the certain site in forest canopy, reaching not only in sides, but also upwards.
Borders are marked by odorous secretions of specific glands of anal area.
Seasonal prevalence in breeding of pterogenettulas is not present, and young
animals may be met at any time of year. Per one year the female brings 2 packs
(4 - 5 cubs in each litter). Young animals differ in darker colouring (light
intervals between dark spots so are small, that colouring seems mesh-looking).
Approximately at four-monthly age young animals start to study “flying” - to
this time the flying membrane at them develops to the full. Sexual maturity
comes at them at the age of 8 – 9 months, and life expectancy may be about
6 years.
Zibetonyx
(Zibetonyx velox)
Order: Carnivores (Carnivora)
Family: Civets (Viverridae)
Habitat: steppes of Central Asia (to the east from Fourseas).
Evolution proceeds by certain principles and laws which were just in the past,
and undoubtfully will be just in the future. Therefore it is quite logical
to expect of recurrence in the future of some phenomena taking place both in
the far past, and within people.
One of such phenomena, convergence with representatives of canine family (Canidae)
and Neocaenic African predator pardinia, is demonstrating by zibetonyx, the
inhabitant of plains in Central and Middle Asia. This gracefully build predatory
animal is the descendant of mongooses (Herpestes) widely settled in Asia. Ecological
plasticity of these small predators has permitted them to occupy temperate
steppes of Asia successfully. And specific conditions of inhabiting - open
plains with high grass - had generated shape of new predator.
Being quickly running predator, zibetonyx has characteristic shape: at it there
are long legs and flexible backbone, small head on rather long and mobile neck.
Jaws of zibetonyx are long and rather weak, its head resembles canine one.
As against cats and pardinia very similar to it, the zibetonyx kills catch
not by one sting: the pack inflicts to the chosen prey numerous wounds and
exhausts it by long pursuit.
At zibetonyx there are sharp sense of smell and good hearing: animal frequently
should hunt in high grass where sight can be practically useless. Ears of zibetonyx
are rounded and wide, but they are located at head sides as at its ancestors.
On the top part of neck the mane especially big at the adult male grows.
Colouring of wool is yellowish with large brown spots merging on sides and
back to short longitudinal dabs. Up to winter (in habitats of zibetonyx snow
does not fall, but weather becomes appreciable more cold in winter) wool becomes
longer and then the mane of males is almost not appreciable.
Zibetonyx is ecological analogue of wolf, but a little bit reduced size. Animals
of this species hunt in packs up to 10 – 15 animals preying approximately goat-sized
herbivores.
The pack of zibetonyxes represents clan with expressed social connections and
hierarchy. Clan is formed of posterity of one parental pair: all females from
packs remain in clan, and maturing males leave it. Females in clan appear connected
by consanguinty and represent two – three generations of one family. Males
come to clan “from the side”, and between them and main pair (clan founders)
there are relations of submission. If friction between males appears strong,
clan can divide at all.
The clan is original “super-organism”: harmonious collective hunting is much
more effective, than single’s one. Zibetonyxes are specialized for hunting
quickly running herbivores: usually harelopes are their catch. However the
important place in menu of zibetonyx is occupied also with small catch: animal
willingly eats lizards, birds and large insects – beetles and grasshoppers.
Main way of hunting is prey chasing. For this purpose zibetonyxes from one
clan stay in line and hide in grass at some distance from each other. Some
individuals (as a rule males are they) drive harelope out from herd and chase
it to animals waiting in ambush. “Passing on” chased animal to each other,
predatorss absolutely not for long time take part in hunting but can “give
all the best” completely, similarly to cheetah. Because these animals combine
hunting tactic of wolf and speed of cheetah it has allowed them to become successful
and prospering plain predators in hunting.
Daily life of clan takes place near natural shelters – small groups of bushes
in which females arrange little lairs and practically in common rear young
growth. Here animals have a rest after hunting. Near shelters zibetonyxes keep
rather carefully and silently, and do not hunt here to not unmask place where
cubs are reared. But borders of clan are plentifully marked by urine and musk
secretions of repugnatorial glands. Usually adult males put marks, placing
them on big stones, bushes and trunks of lonely growing trees.
Cubs (2 - 3 in pack) are born blind and covered by short wool. Their colouring
is very dark: brown, paws and ears colored black. For the fourth day of life
at them eyes open and then they start to hear. And ten-day cubs already try
to walk. Only first two weeks of life young zibetonyx knows exclusively care
of native mother. Growing up, it becomes object of gentle care at once of several
females of clan, including old ones which can not bear posterity any more.
They play role of “baby-sitters” when clan goes hunting. Such feature raises
chances of survival even at deserted cubs. One female is capable to bear posterity
twice per year.
At the age of four months young animals already change youthful dark colouring
to spotty adult one. They study to hunt about two months, and then when females
of clan have new cubs, young males are driven away. They can live vagrant life
on border of territories of several clans, satisfying by remains of their catch.
Frequently young males form independent packs try to drive out adult males
from any not numerous clans, or to drive off young females from near clans.
Usually bachelors are much more aggressive than males have formed pair. They
easily get involved in fights though they can suffer serious loss from strong
clans: females protecting posterity can fight up to death. Usually many young
males perish in such skirmishes, and sex ratio in generated and breeding clan
is 1 male for 2 – 4 females.
Sexual maturity at females takes place at second year of life, at males – at
third year. Life expectancy is about 15 – 18 years.
Mauritian paralutra (Paralutra nana)
Order: Predatory (Carnivora)
Family: Civets (Viverridae)
Habitat: rivers of Mauritius Island.
The fauna of Mauritius Island had strongly suffered from people influence.
Many species of animals representing groups which could not appear on island
by natural way had been introduced to the island in different times. These
animals were monkeys, rodents, various predators and domestic hoofed mammals.
Such inflow of species new to the ecosystem of the island had taken place in
very short term according evolutionary measures – within several centuries.
As a result the native fauna and flora of island had degraded. People could
interfere with influence of introduced species to the endemic nature of island
with the help of nature protection actions till some time. But after the disappearance
of mankind the rests of island primordial nature were destroyed by introduced
species. In Neocene flora and fauna of Mauritius are presented mainly by descendants
of introduced species had formed the complete and balanced ecosystem. In most
cases inhabitants of islands differ in diminutiveness, and only few species
of reptiles and birds are the exception. Descendants of introduced species
had occupied all ecological niches were given to them by the nature.
Rivers of island are shallow and short. Descendants of the fishes introduced
by people – true gourami (Osphronemus goramy) and goldfish (Carassius auratus
auratus) live here. Occasionally sea fishes swim into the rivers of island,
and there are some species of river shrimps and crabs. These species make the
basis of food pyramid at which top there is one endemic species of predators
adapted to life in water – the Mauritian paralutra. This animal is the descendant
of mongooses (Herpestes) introduced to the island and perfectly swimming predator.
Paralutra eats any water animals which can be caught, and supplements a diet
with tubers and sappy leaf petioles of water plants.
Mauritian paralutra is a small animal. This is the natural phenomenon in the
world of small island, where resources and vital space are strictly limited.
Length of adult animal is about 60 cm including long tail. By appearance and
constitution paralutra resembles true otters (Lutra) of Holocene epoch: this
is a short-legged animal with flexible body and membranes between toes. Fingers
of forepaws of paralutra are not connected by membrane. They are sensitive
and mobile, and have small claws. With their help paralutra searches in thickets
of plants for shrimps and small fish. On the coast paralutra clears and combs
fur by fingers. The wool of animal has grey color, on throat there is white
spot, and cheeks are also white.
Head of paralutra is flattened, with short wide nose and long postorbital part
of skull. Eyes are shifted to the forward part of head and upwards, therefore
animal can breathe and look over vicinities, having put out from water only
a part of head. Above the nose of animal there is a site of hairless skin with
which help paralutra digs bottom in searches of crustaceans and roots of plants.
External auricles at paralutra are lack, and the ear aperture at diving animal
is closed by ring muscle. Tail of paralutra is long, flattened, covered with
naked skin. At movement under water animal presses forepaws to body, and swims,
wavy beding by all trunk. For slow and maneuverable swimming among underwater
thickets animal uses hinder legs.
Paralutra eats small fish, not attacking large fishes. Hunting animals try
to push aside school of fishes to the bank and attack them in common. Also
this animal eats shrimps and crabs, and ravages nests of coastal birds.
Musk glands located under tail produce plentiful oily secret with which help
paralutra makes its fur water-repellent. To not frighten off fish, this product
of secretion till the process of evolution had lost smell that is completely
atypical for civet. This mammal smears secretions of musk glands on forepaws,
and pounds secretions on fur, carefully rubring them in hairs and combing them
by claws. Ritual of mutual fur caring helps to support friendly relations between
the representatives of species living together.
This is sociable mobile animal. Paralutras settle on steep riverbanks by whole
families including some females connected by consanguinty, and males not related
to them. The family of these animals digs out some inhabited holes, one of
which is occupied with main pair. These animals breed the year round. The female
can bring posterity two times per one year, 3 – 4 cubs in one litter. Preparing
to birth of posterity, female deepens the hole and digs out wider inhabited
chamber, rather than usually does. Cubs are born blind and deaf, covered with
short dark wool. They start to move in hole at the age of about 12 days, and
monthly cubs already are able to swim as well, as adults. At the age of 3 months
young animal becomes completely independent; half-year old female already gives
rise to posterity.
Paralutras constantly support contact with each other with the help of voice.
They can utter various sounds – they chirr, peep and whistle. At danger animals
yelp like small dog. After feeding animals gather and clean wool to each other.
At the rest paralutras are cautious and prefer to gather at shallows and small
islands in the middle of the river where it is impossible to creep to them
imperceptibly. At suspicious sounds and sudden occurrence of large animals
at the riverbank they disappear under water. Main enemies of paralutras are
maurosmiluses – small saber-toothed ground predators of feline family; in water
some sea fishes swimming to rivers of island can attack these animals.
Grumbling
ant-mungo (Myrmungopsis myrmivorus)
Order: Carnivores (Carnivora)
Family: Ant-mungoes (Myrmivoridae)
Habitat: Atlas Mountains, Middle East, Southern Europe (coast line of Mediterranean
swamps)
Some species of live creatures have success in struggle for existence due to
development of habitats and food resources inaccessible to other species. This
mammal species belongs to such animals. It is the representative of new family
of Neocaenic mammal, eating exclusively social insects. The ant-mungoes family
had descended from civet family: some small species of family had specialized
in feeding by insects, and further this specialization has resulted in occurrence
of new group of animals. Suricata (Suricata), social species of small civets
became direct ancestor of representatives of present family.
The ant-mungo during evolution has got common for insectivorous animals appearance
features: at this animal there are wedge-shaped extended head, small mouth,
reduced lower jaw and teeth. The basic feeding organ is tongue, capable to
extend forward approximately for one and half lengths of head. Nostrils are
located on tip of muzzle, during the feeding process they close and animal
holds the breath. Claws on ant-mungo forepaws are long, during movement the
animal turns them in and supports on the back side of hand similarly to anteater
or gorilla (in these places “callouses” of dense thick skin develop). Claws
are used for digging insect nests and for protection against enemies. The wool
of ant-mungo is short, dense and rigid, bristle-like. It protects animal from
stings of insects. Besides animal has significant immunity to insect poison.
Colouring of wool is dingy: monochromic gray-brown, stomach is light grey,
legs and tail are darker. Forepaws are shorter then rear legs; at movement
the back of animal is inclined forward. Digging out an ant hill or termitary
ant-mungo can stand on hinder legs for a long time.
The grumbling ant-mungo reaches length 40 cm (tail is up to 30 cm) and weighs
about 5 kgs.
Similarly to the ancestor, ant-mungo has kept social habit of life differing
in it from other Neocaenic “anteaters”. This species lives in families of 4
- 7 animals (breeding pair of adult animals and cubs of the current year).
Family occupies rather extensive territory with thickets of bushes and light
forest where many nests of termites and ants are located. Except for them animals
eat small snails, swallowing them entirely with shell, and also larvae of beetles
and caterpillars of butterflies. Animals communicate among themselves with
the help of sounds similar to rumbling and grunt.
This animal lives in hot, almost waterless areas, therefore it shows greatest
activity in twilight. The group of ant-mungoes spends day in temporary hole
dug on the eve on morning, and at the sunset leaves it for feeding. In the
evening animals eat mainly ants gathering in ant hill for the night, at night
– by termites coming on surface, and gather snails crept out on plants covered
with dew in the morning, or search for beetle larvae under stones and tree
bark.
Pairing of ant-mungoes occurs approximately for two months before rain season,
and cubs are born just in this season. It is justified biologically: milk producing
at the female takes plenty of water, and in rain season it is easier to fill
up internal water stocks. Once a year, in the beginning of rain season female
gives rise to five - eight cubs. They are born blind and deaf, female hides
them in specially dug hole. She feeds posterity, coming back in hole some times
for night. At this time male shows original care of her: he gorges on for future
use, and then in addition feeds up the female, belching food to her mouth.
At one month after birth the young growth already can follow adult animals,
and the female actively trains posterity to search for food. Usually pack stays
with mother up to the following season of pairing when the parental pair banishes
posterity from the territory. The young growth reaches sexual maturity at half-year
age, and life expectancy is about 8 years. Main enemies of grumbling ant-mungoes
are predatory mammals and snakes, but most of all these animals perishes during
long droughts.
Awfulest marafil (Marafil perhorridus)
Order: Carnivors (Carnivora)
Family: Hyenas (Hyaenidae)
Habitat: South Africa, mountains and plains.
In Neocene South Africa appeared separated from Holarctic zoogeographic area
by the zone of impassable tropical woods. They had appeared when East Africa
(Zinj Land) had separated from the main part of the continent, and the area
of dry climate had considerably reduced because of evaporation of water from
Tanganyica Passage.
The majority of groups of herbivores appeared in Neocene had not got here;
therefore Southern Africa became one of places where many representatives of
hoofed mammal fauna characteristic for Ethiopic zoogeographic area had kept.
In South Africa in savanna descendants of
water chevrotains similar to deer
wander, and tall descendants of dwarf duiker
antelopes similar to primitive
giraffes browse foliage of trees. Representatives of animal kingdom of new
epoch – huge descendants of pigs armed with massive bone outgrowths on head
– also are found here. By the efficiency ecosystem of Southern Africa does
not concede to North African savannas of early historical epoch.
The abundance of catch involves various predators, and relative stability of
this region had allowed to some representatives of Holocene fauna of predators
to be kept in South Africa. Among civets and cats of South Africa one local
predator looks separated – it is the last representative of hyenas’ family.
This family had been widely distributed in Eurasia in Holocene. Till the ice
age some of its representatives had wandered in snow-covered woods of Europe,
tolerating severe winters. But after mass extinction of hoofed mammals in the
areas mastered by people the area of family was sharply reduced. In historical
epoch the family was submitted only by five species, one of which was extremely
rare.
Awfulest marafil (“marafil” is the Arabian name of hyaena), the descendant
of spotted hyena (Crocuta crocuta) widely settled in past, lives in South Africa
This animal represents the deification of “predator” concept, doubtlessly occupying
the top of food pyramid in local ecosystems.
Awfulest marafil is the largest species of hyenas among ever dwelt on Earth,
and, obviously, it is the largest representative of the carnivore order in
Neocene. Growth of the adult female at a shoulder reaches two meters at weight
about 700 kgs. Male is smaller, rather than female: it weighs about 500 kgs
and differs in lighter constitution. By the appearance marafil resembles fossil
Arctodus bear dwelt in Pleistocene at the territory of North America. The female
of marafil differs in sloping back (back legs are shorter a little, rather
than front ones) and wide chest. Forepaws at it are very muscular – with their
help the animal knocks prey off feet. Feet at the female of marafil are rather
wide due to what animal can chase prey on marshy ground. At the animal there
is short tail with hairy brush of lengthened hair on the tip.
At marafil there is short head with powerful high jaws. Canines are rather
short; molars and premolars of animal work as crushing tool – marafil female
can shatter the backbone of rhinoceros-sized prey by one bite.
The skin of marafil female is covered with rough short wool of brown color.
There are vertical strips of black color on shoulders and sides and some black
spots on hips (sometimes they may be absent). Wool on head of animal is black,
but around of mouth there is a strip of white wool – this attribute has the
important value at the communication of animals. Marafils, especially females
of this species, are rather aggressive animals, and in many cases it is vital
for them to be able to distinguish mood of the congener from afar to behave
in appropriate way.
Very sharp sexual dimorphism is characteristic for marafils. It is shown not
only in difference of body sizes, but even in colouring. Skin of marafil male
is grey with numerous black spots merging on shoulders to vertical strips.
On the neck of male there is a mane of long grey hair. Males look more harmonous
and long-legged, rather than females and can run much faster. Also at the male
there is longer and narrow muzzle with more advanced canines which partly jut
out from closed mouth.
Males live and hunt in packs of ten individuals, living mainly at the boundary
of territories of female clans. They do not make constant refuges, and spend
the night in the same place no more than two nights in succession. Females
of marafils, on the contrary, make constant lairs in bush and come back there
every day. They lodge in clans of 4 – 5 individuals connected by consanguinty
(as a rule, it is the female and her senior daughters). The number of marafil
males in population is smaller, than the number of females: it is connected
with the high mortality level of these animals at young age. When young marafils
grow up, young females dominate over males compelled to be content with the
rests of their catch. When the young marafil male reaches about one-year-old
age, it is expelled from clan, and some time it leads single life, eating the
scraps had remained after hunting of parental clan. Groups of marafil males
can unite, forming male packs.
Sexual dimorphism of marafils is shown also in hunting preferences: there are
different catch and different hunting tactics at males and females. Females
chase mainly sluggish catch – huge bullhogs. Males prefer to hunt swift-footed
hoofed mammals, badgering them by pack like wolves. Hunting habits at different
genders also differ – females hunt at night, attacking resting and feeding
animals, and use during hunting sharp sense of smell. Males hunt in day time
with the help of sight.
Marafil females being in group are very aggressive to neighbours. It happens
they can attack males of their own species, kill and eat them as if other prey.
Therefore pairing at marafils occurs outside of territory of female clan. For
pairing males mark with urine objects well appreciable from afar (for example,
lonely growing tree, large stone or termitary) at the edge of territory of
females, and wait for female near this place. Thus they drive other males off
from the marked sign. Also males leave some of labels at the territory where
the female clan lives. Usually they do at the afternoon before females will
leave on hunting. But at the same time males try to keep together and to be
on alert. They succeed to have a sleep only in second half of night and in
the morning approximately till midday when females come back home from hunting.
The female ready to pairing leaves clan, finds the site marked by males, and
males pair with it by turns. Before pairing the female shows absence of aggression
to males: she lowers head and presses ears. Males showing the superiority run
near her by whole pack with lifted heads and occasionally bite the female to
shoulders and groats.
Similarly to all large animals, marafils breed seldom: each female gives rise
to posterity alternate years. Pregnancy lasts about half-year, and cubs are
born to the beginning of rain season. They suck milk till four – five months,
but start to try meat food at the third month of life. In litter at one marafil
female it happens three – four cubs. Usually only half of posterity survives
up to independence.
Young females become able to breeding at the age of 4 years, and males since
3 years. Life expectancy at females of this species reaches 50 years and more,
and males seldom live up to 40 years.
Siberian
sabertooth (Machairolynx sibirica)
Order: Carnivores (Carnivora)
Family: Cats (Felidae)
Habitat: Northern Eurasia, Beringian isthmus, northwest of Northern America.
After mass extinction at the boundary of Holocene and Neocene the variety of
predatory animals has appreciably decreased, and some their families have
almost completely disappeared. The most successful among them representatives
of weasel and civet families appeared, but except for them in separate areas
of Earth representatives of cats (Felidae), the most specialized family
of predators, and also some other predatory animals were kept. Large cats have
died out, but the family has escaped due to the variety and ecological plasticity
of small cats which were developed later to large forms. One of the largest
cats of Neocene is Siberian sabertooth, the descendant of lynx (Lynx).
This relic representative of feline family is the large prey hunter. It is
possible to guess it easily, having looked at the constitution of animal -
this predator is not similar to cheetah or leopard chasing swift-footed catch.
Body of sabertooth is very massive: this is a tiger-sized animal (weighting
about 350 kgs), but its figure resembles bear more: legs are thicker with wide
pads (animal began almost plantigrade one). Back legs are a little bit longer
than front ones, back is inclined forward. The short tail has got to this animal
as the inheritance from the ancestor. Wool colouring is sandy-yellow with dark-grey
spots, to winter it changes to white with ashy shade (this phenomenon is the
exception among cats not changing color of wool in winter), stains on winter
fur completely disappear. The winter wool is much longer and also thicker than
summer one: it is the adaptation to life in conditions of continental climate
of Northeast Asia with rigorous snow winters.
The large prey hunter, Siberian sabertooth is perfectly equipped for preying
of large thick-skinned animals. Its head is rather large (length of skull is
up to 30 cm), the bottom jaw is very mobile and can open widely. In the top
jaw long 15-centimetric saber-like canines with the wide bases stick up – it
is the main hunting weapon of animal. More successfully to use this weapon,
the skull is in addition “strengthened” – bases of canines are very wide, skull
is high with short obverse department. Besides at sabertooth there are very
strong neck and powerful forepaws – it is directly connected to way of hunting
of this animal.
Siberian sabertooth is the tireless nomade of open spaces of Siberia and Beringia.
This animal literally “pastures” herds of large herbivores, following them
during migrations. The most important catch of an animal is obda,
huge herbivorous animal of Siberia. For hunting for this giant sabertooth uses
tactics of attack from ambush, but it frequently avowedly attacks animals kept
up with herd. Predator attacks catch sideways, striking during the throw by
one paw on shoulder, by another one – on head of prey. Usually young or weak
animal falls aside, and it is easy for finishing off by bite in throat. At
the sting canines of sabertooth pass below backbone (in other way they simply
can be broken) and break off blood vessels and trachea of prey. If the chosen
prey is strong enough, predator simply wounds it by bite and follows herd,
expecting while wounded prey will get behind neighbours, bleeding profusely.
One carcass of obda suffices to the predator in summer time for one week -
while meat will deteriorate to the uneatable condition. In winter when meat
is better kept, the preied animal suffices approximately for two weeks. But
more often such catch should be divided with set of feathered and four-footed
scavengers; therefore sabertooth had to hunt more often. One animal theoretically
should supervise territory 200 - 300 square kilometers, but it leads nomadic
life, therefore sometimes “shepherds” of different obda herds meet on summer
pastures of these animals. At this time pairing and birth of cubs concepted
the last year happen. At summer obda pastures sabertoothes sometimes unite
for hunting, together getting the next dinner.
Pairing of sabertoothes occurs at the end of summer shortly before leaving
of obda herds to the south. The female ready to pairing calls the male by sounds
similar to loud bass mewing. Usually near such female two - three males keep
but only choosed by female one can couple with her. Between males there are
skirmishes during which contenders beat each other by paws and show force,
peeling bark from trees. Males ready to pairing also mark by urine tree trunks
– smell plays the important role for the female in choice of the partner. When
pair is formed, male and female banish single males together. Pairing at these
cats repeats many times during three - four days while the female is ready
to accept attention of the male. At this time the male looks after her, licking
her wool. He also marks a site of wood around female with urine to warn contenders,
that the present female is occupied. During courtship games animals eat of
nothing. After pairing the female walks out from the territory marked by male.
Pregnancy at this species proceeds a long time: more than nine months. The
reason of it is that fertilized oosperm does not develop at all about half-year.
Development of embryos begins approximately at first half of winter, and actually
proceeds little more than three months. Cubs are born just when the female
comes to the north after wintering in woods.
For birth of cubs the female sabertooth makes a lair under roots of trees or
among stones. In pack there are two or three cubs. Newborn sabertoothes are
covered with dark-brown wool on which stains are already appreciable. At them
eyes and ears are closed, and first two weeks of life they are absolutely helpless
and completely depend on mother.
Within four months, before autumn obda migration cubs grow and develop quickly.
At this time they actively study to hunt, accompanying with mother. During
migrations behind obda herds sabertooth cubs remember district, and later,
becoming independent, begin “shepherd” herds.
Young growth stays with mother up to one-year-old age. When families come to
northern obda pastures again, young animals leave mother and start to hunt
independently. Sexual maturity at young females comes at the second year of
life, at males at the third year. Life expectancy can reach 40 years.
In east areas of Northern America Siberian sabertooth is replaced by close
species of easier constitution – Nearctic
sabertooth (Machairolynx nearctica), or missopeho (named
after giant lynx of the North, the hero of Indian folk tales). At this animal
canines are shorter; colouring is lighter and does not vary for winter. Constitution
of Nearctic sabertooth is more graceful: legs are longer, size of body is less.
It is connected with specialization to feeding by smaller swift-footed prey.
As against to Siberian sabertooth, these species is settled and territorial.
Awful daggerclawer (Pugionyx deimus)
Order Carnivores (Carnivora)
Family Cats (Felidae)
Habitat: Eastern Asia, Far East (area of moderate and subtropical climate),
flatland woods, foothills.
In Holocene the cats family has suffered the big damage from human activity.
Number of large cats was sharply reduced, and their areas have catastrophically
decreased. Small cats have suffered from hunting for the sake of valuable fur.
The unique representative of family has received the big advantages in a survival
in human epoch is the domestic cat Felis catus. Due to people this predator
was settled practically in whole world. It became a stimul for evolution of
the present species. As against the majority of domestic animals dependent
on people, the domestic cat easily runs wild. Therefore after disappearance
of people this species have not died out in common with them, and have continued
to exist. The majority of Neocene felines descend from several species of rather
small cats have survived hunting and ecological crisis.
Cats are successful predators; therefore descendants of domestic cats have
taken the high place appropriate to their abilities in food pyramids of new
biosphere. And in Neocene among them large animals from the number of the most
terrible land predators of Earth have appeared. One of them is the awful
daggerclawer, specialized large prey hunter.
It is very large cat: it is lion-sized, but it seems larger because of longer
legs. At daggerclawers characteristic features of appearance, common for cats,
were kept: rather short jaws, large canines, binocular sight. Daggerclawers
are solitary animals.
The body of daggerclawers (in the Neocene world there are few species of these
cats) is covered with fluffy wool. At the awful daggerclawer the colour of
wool is similar to “tortoise-shell” color of some domestic cats - on grey background
there are big, almost symmetric dark spots. Tail of animal is long, on it spots
merge to non-uniform cross strips. A tip of tail of animal is white. White
sites are on the bottom part of muzzle; throat and breast are also white. On
hinder legs of animals of this species occasionally there are white “stockings”.
Forepaws of daggerclawers are colored contrastly and brightly: from an elbow
up to fingers they are covered with alternating wide black and white strips.
This colouring is the important element in behaviour of animals as it is used
in rituals of an establishment of domination. It is shown only at adult animal
of both sexes. Not sexual matured animals have dark forepaws.
At capture of catch cats actively use forepaws. And this feature of behaviour
has received development at the present species and has reached high specialization.
On paws of cats there are five fingers from which four ones base on the ground,
and the first, the shortest one, does not touch ground. At daggerclawers on
first fingers of forepaws crescent claws of huge size (its length is up to
20 cm) were developed. In rest this claw is pressed to the paw and its end
is directed back, not preventing walking and running. On the internal sides
of wrists of the animal there are sites of very thick cornificate skin, and
this claw adjoins directly to this one. Also this claw is partly retracted
in skin cover. The “fighting” claw of the daggerclawer is partly similar to
claws of small predatory dinosaurs like Deinonychus and Velociraptor.
The daggerclawer eats large ground animals. Attacking prey, it at first creeps
to it to probably closer distance, and then rushes to the planned victim. Due
to long legs it can accelerate momentum up to 60 kms per hour on the short
distance. At the moment of solving jump the animal shows its “fighting” claws,
and puts to the victim deep wounds by them. When impact has falled on the neck,
catch of the daggerclawer, even having escaped from its claws, quickly bleeds
profusely and perishes. In case of successful attack the daggerclawer, except
for impact by claws, delivers to prey sting in neck or in the basis of skull:
this is the “inheritance” stayed from its ancestor, the domestic cat.
All daggerclawers are solitary territorial animals. Usually at them as at tigers,
the territory of the male includes territories of several females. Animals
mark territory, fleecing bark from trees, and rubring the bared wood with a
secret of chin repugnatorial glands. Males also mark the territory by urine.
During an establishment of domination relations animals pay special attention
to colouring of forepaws of the contender: the more contrast coloring, the
animal having it is stronger. At young animal paws are dark; at old ones they
are “gray-haired”: strips become not so contrast. Duels for territory and place
in hierarchy, as a rule, are limited by demonstration of “stretching” of contenders:
thus animals by turns bend backs and extend forward forepaws, showing their
colouring. “Stretching”, each animal tears off by claws a part of grass, breaking
through deep furrow by “fighting” claw. Such ritual behaviour helps to avoid
fight, and contenders can estimate force each other. It is accompanied by loud
mewing and squeal.
Pairing occurs in middle of winter, kittens (2 - 4) are born in first half
of spring. They are covered with dark wool, blind and deaf. The female feeds
with them with milk about four months, and then even more half-year young animals
keep with mother. They abandon her in winter, and more often they are banished
by males during a rut.
Sexual maturity comes approximately at three-year age. At this time at males
strips on forepaws start to appear. But most successfully males are coupled
approximately since the fifth year of life: females give obvious preference
to males with striped forepaws.
Life expectancy of the awful daggerclawer reaches 30 years.
There are some other species of daggerclawers distinguished in smaller size
in East Asia in moderate and subtropical latitudes:
Fisher daggerclawer (Pugionyx piscivorus) is a predator up to half meter at
a shoulder. Length of its body is about one meter; a tail is about 60 cm. The
head is dark with white area on the tip of muzzle and throat, above eyes there
are white spots; wool has striped colouring. Forepaws up to an elbow are white,
occasionally on them there are one - two cross strips. The claw is rather larger,
than at the awful daggerclawer: its length is about 15 cm. In rest it is strongly
unbend back, resembling cock heel.
The animal inhabits marshlands, riverbanks and lakes. The fisher daggerclawer
willingly swims and prefers to not leave water far. The animal eats fish and
aquatic animals: turtles, frogs, waterfowl and rodents.
Bird-catching daggerclawer (P. saltatus) is the smallst
species of genus (length of body not including tail is about 60 cm; tail is
about half meter
long). It is settled more widely than other species: this cat inhabits areas
from plains up to mountain woods. It also differs in especial addition: hinder
legs are strong, appreciably longer than forepaws (back is inclined forward).
Colouring of wool is spotty: on background from sand up to red there are dark
brown spots. Forepaws armed with sharp crescent claws are colored black with
white spots. This animal eats mainly birds. It dexterously forces down flying
up birds, killing for one jump up to five ones from flight. It eats catch,
keeping it in forepaws and sitting on back legs (by same habits Holocene caracal
(Felis caracal) had differed).
The idea of the present animals is prompted by Simon, the participant of forum.
Tiger raptor cat (Felinoraptor tigrinus)
Order: Carnivors (Carnivora)
Family: Felids (Felidae)
Habitat: South America, subtropical and temperate areas; light forests and
woods in foothills.
At the boundary of Holocene and Neocene human activity had caused the mass
extinction of large herbivores. It had resulted in extinction of set of predators,
including large cats of genera Panthera (the majority of large cats), Uncia
(snow leopard), Acinonyx (cheetah). In same time many smaller cats became the
extremely rare as a result of hunting or destruction of natural environment.
However, the felid family had survived in Neocene. But in most cases it is
the consequence of worldwide settling of domestic cats which descendants had
run wild and had taken part in formation of new ecosystems in various parts
of the world.
In Neocene connection of South America with other continents had interrupted
again. Some time in early Neocene the Antilles land bridge existed, and using
it land animals could settle, but then it had disappeared. In conditions of
isolation descendants of puma (Felis concolor) had turned to very large predators
of frightening appearance. These cats of South America represent the unite
group which has the general distinctive attribute – the original adaptation
for killing of large prey. The inner toe on hind leg of these cats had strongly
changed: it became little bit shorter than the others, but on it the huge strong
claw is developed. In rest it is retracted into special sac formed by skin
plica. When these cats attack prey, they seize it by forepaws and tear prey’s
flesh between ribs or on the stomach by “battle” claw of rear legs. This adaptation
resembles very much the feature of hind legs anatomy of small theropod dinosaurs
of Cretaceous, and hence the name of such predators – raptor cats. It is the
original experiment of evolution: in South America the kind of contrast to
Asian daggerclawers, massive cats with similar “battle” claw developed on forepaw
had evolved.
The constitution of these predators is rather light and “universal”, permitting
them to swarm up trees and to move on rocks good. Due to this feature raptor
cats are widely settled in South America in places where enough plenty of large
herbivores lives. They are the specialized hunters on large sluggish extraction
- large rodents barocavia and giant paca. Occasionally they attack on young
tapirotheres and groundsloth rodents. These cats attack prey from an ambush,
strike it a blow by “battle” claws, and jump off aside. If the animal had not
died from the first attack, the cat pursues prey, not repeating attack while
the victim will die from loss of blood.
In South America three well differing species of these large cats live. Largest
of them is tiger raptor cat – the lion-sized predator. This species has dark-rusty
skin with numerous vertical strips of black color on body and paws. Ears and
top part of head of this cat are black, and cheeks and throat are white. Tiger
raptor cat inhabits flat districts and foothills, preferring the mosaic landscape
where areas of grasslands, woods and rivers are combined. It is a solitary
predator; each individual occupies extensive territory up to 30 square kilometers
in area. Despite of large size, tiger raptor cat is freely able to swarm up
trees and rocks and also to swim. It easily attacks prey in water, and its
prey may often become barocavia or giant paca – semi-aquatic rodents. Occasionally
these cats catch young thyranncharaxes - huge predatory characid fishes.
This species of cats marks borders of territory by urine, leaving it on bushes,
tree trunks and termitaries. The territory of the male substantially overlaps
territories of several females living in the neighbourhood. Usually females
concern tolerantly enough to the presence of male out of breeding season, though
the female with cubs or near the prey can be aggressive to him. Males concern
to each other much more aggressively, and at the meeting necessarily display
the force to each other. This demonstration of force at well armed predators
passes very originally, practically without the direct contact: they sort out
their relationships in original competition, by turns or nearly so simultaneously
jumping on tree trunk. Having seized claws in bark, males hang on a trunk till
some seconds, hissing against each other, and then jump off. Usually the winner
is the one jumping higher and keeping on tree for longer time. If males are
equal in force, they can combat. Also the adult male concerns aggressively
to male youngsters; the majority of them perish till first months of independent
life from paws of adult males. In rare cases the old male can become cannibal
and steal cubs at females living in the neighbourhood.
Courting the female, the male leaves marks in her territory to give the female
the opportunity to get accustom gradually to his presence. “Unready” female
can rush on the male if he will appear too suddenly – usually it happens so
at young males. The female is ready to pairing till several days, and all this
time the male preserves her against other males, and frequently pairs with
her. When the rut stops, female banishes the male, showing aggression to him.
After six-monthly pregnancy, in the beginning of rain season female gives rise
to the litter of from two to eight cubs. In big litter usually only three or
four cubs have an opportunity to grow up to independence. For rearing of posterity
the female arranges the den in bush or in wide hole dug out additionally to
the necessary size. Cubs are born blind and deaf. Their wool has the same strips
as at adult individuals, but strips are rather wide; therefore cubs seem darker,
than adult individuals. At three-monthly age they leave a den and start to
study to hunt actively. Young animals spend with mother about one and a half
years, therefore the female brings posterity only once in two years. Young
animals become sexually mature at the age of three years.
The close species lives in mountains of South America: the Andean raptor
cat (Felinoraptor andinus). It is little bit lower growth, than tiger raptor cat,
but is built much stronger and heavier than it. The “battle” claw on hind legs
at this species is advanced not so extremely – in mountains there is less number
of large prey, and Andean raptor cat eats mostly smaller animals – birds and
rodents. But it also hunts young mountain tapirotheres.
One more species, the Chilean, or western raptor cat
(Felinoraptor occidentalis),
lives in narrow land strip between Andes and Pacific coast of South America.
It is settled from equatorial latitudes up to Tierra del Fuego. This species
of cats differs in pale yellow skin with some dark spots. Northern populations
living in warm latitudes have more gracile constitution and wider ears, and
southern ones, living in areas of cool climate, are smaller and short-legged,
with almost one-colour rich fur. In the extreme south of South America this
species of cats is superseded by larger Andean raptor cat. Chilean raptor cat
almost does not live in mountains, preferring woods in foothills and in plain
areas. These predators search for food even at the coastal strip, and frequently
eat bodies of dead algocetids cast ashore during the storm.
Banshi
saki (Neomyrikina banshi)
Order: Primates (Primates)
Family: Cebids (Cebidae)
Habitat: Amason region, rainforests and woods in foothills.
On boundary of Holocene and Neocene big tests for durability expected the primates
order. The majority of representatives of this order in the daily life is strictly
connected with woods. Here they find shelters and food. And when on boundary
of Holocene and Neocene the global congelation began, the majority of primates
has died out. In New World in Holocene this group was submitted by three families
which number and variety were fairly undermined by human activity. Therefore
chances of survival at most part of species were minimal. But nevertheless,
among these animals some ones were to found has gone through global ecological
crisis and has successfully occupied Neocaenic woods.
When night goes down on rainforests of Amason region, it is possible to see
some representatives of Neocaenic monkeys of New World. In moonlight in underbrush
small groups of night monkeys banshi saki wander. These animals are rather
large for monkeys of New World: length of body of adult individual is up to
50 cm not including long tail. They are colored rather dimly in comparison
with monkeys of Holocene epoch. At these primates head is covered with white
wool, on neck, cheeks and crown long white hair forming original “pallium”
grow. Very big brown eyes with round pupils indicate that it is nocturnal animal.
In addition eyes are “penciled” from above by black wool. Only on chin males
and females have red beard (at males it is much longer, than at females). Males
in addition are decorated with rich white moustaches which are hanging down
downwards from corners of mouth. Bodies of these monkeys are covered with dense
grey wool, on chest and stomach wool is darker. The tail tip is white, but
set off by black ring.
Jaws of these monkeys are short; at males large canines slightly jutting out
from closed mouth are advanced. Nostrils as at all wide-nosed monkeys are bordered
by wide nasal sept. The forward part of muzzle at the adult banshi saki is
covered with thin wool, and at old animals it may be hairless at all. Ears
are short, rounded; only their tips jut out from wool.
Hands and feet at banshi saki are hairless, covered with grayish-pink skin.
The thumb at these monkeys does not oppose to others (this feature distinguishes
monkeys of New World from other monkeys).
Banshi saki is one of the cleverest inhabitants of Amazon woods. At them there
are rather large brain and complex behaviour including set of acquired skills,
carefully kept in collective experience. Monkeys keep contact with each other
uttering various sounds: they howl, chirp, smack and squeal silently. In total
in their “dictionary” there are some tens of sounds and their combinations.
Besides they watch white tips of tails of the neighbours, flashing in darkness.
These monkeys distinguish colors rather badly, but their night sight is very
keen.
Groups of these original primates amount up to 20 individuals, and they are
headed with dominant pairs: all males submit to the male, the female bosses
over all females of group. If the dominant male chooses to itself the female
not belonging to number of animals of the supreme rank (at these monkeys pair
is forming according personal sympathies), the group status of “Cinderella”
automatically grows.
These monkeys wander in forest, finding drop fruits, insects and small vertebrates
on the ground. But it is rather dangerous to make it: at night numerous predators
wake up and go to hunt. Usually these monkeys are very cautious, and some individuals
of medium rank are sentinels of group. Such duty allows them to lift the social
status though prevents to search for food normally. Feeling approaching of
any predator sentinels notify group about it by sharp squeal. Usually monkeys
try to save from predators on trees. If they can not make it for any reasons
they play awful performance for which have received the name.
Frightening off the enemy, banshi saki spread manes asides, having turned by
faces to the enemy. Their heads turn to terrible masks: stains above their
eyes look on the background of white mane as huge eyes, and spreaded beard
seems an aperture of huge opened mouth. Moustaches of males besides create
impression of canines in “mouth”. This performance is accompanied by loud shrill
squeal (in the English folklore banshies are creatures remarkable in heart-breaking
voices). To seem larger monkeys rise on hinder legs. Then the white mane is
even more brightly appreciable on the background of black stomach. Usually
not any predator endures monkey’s “performance” for a long time, preferring
to run away. But even if its hunting appears successful and one of monkeys
becomes its prey, banshi sakis all the same cry at the predator by all group,
chasing it forthwith – it can secure them against the following attack.
Once a year the female gives rise to two cubs. They at once seize wool on her
back, and first two weeks of life they travel on mother. Gradually young monkeys
start to master the world, to try food of adult animals and to play with neighbours.
Half year old animals already move independently, and one-year-old ones live
independent life. At the age of two years young animals can have posterity.
Life expectancy of banshi saki reaches 15 years.
South-American barbudo (Barbudo papioformis)
Order: Primates (Primates)
Family: Cebids (Cebidae)
Habitat: north of South America, forests and bush in foothills.
Primates had appeared in South America in early Oligocene, and during their
natural history were represented by exclusively arboreal forms having greater
or lesser degree of specialization to tree-climbing. Some species of South
American monkeys had evolved tenacious tail allowing to cling to branches and
to hang on it. All these monkeys form a group of New World monkeys, which prominent
feature were very widely placed nostrils.
In Holocene epoch many kinds of New World monkeys were destroyed as a result
of hunting, catching for zoos and deforestation of Amazon basin. The most specialize
arboreal and local endemic species had especially strongly suffered. But some
species managed to survive, to go through climatic changes at the boundary
of Holocene and Neocene, and to inhabit again forests of South America, which
had gradually started to restore in early Neocene. Common squirrel monkey (Saimiri
sciureus) became the most successful species of South American monkeys. Its
success had been determined by a number of factors: they easily adapted to
life in broken forests and bush, frequently went down and could easily move
on the ground, were numerous and differed in small size. The majority of monkeys
of Neocene South America descends from Saimiri. One of these species is large
South American barbudo.
Barbudo are medium-sized monkeys (body length is about 40 cm) having rather
massive constitution. They live in big groups on the ground in forests of various
types in central and northern part of South America. These monkeys meet especially
frequently in foothills and among bushes, where they find a lot of food. They
are able to climb on trees, but do it reluctantly, because they are more adapted
to life on the ground. Young individuals climb on trees more often.
Barbudo has massive trunk with voluminous stomach. Front and hind legs are
of approximately equal length, therefore the back of animal is slightly inclined
back. Tail of barbudo is very short (about 10 centimeters long); with its help
monkey can only express the mood, and it does not take any part in movement
on trees.
Head of barbudo is rounded and has short strong jaws. Males have large canines;
they frighten contenders and defend against predators with their help. Females
have lesser canines, and during the repulse of predator attacks they keep near
cubs in rear of the group while all males of the group in common attack predator,
uttering loud cries.
The body of barbudo is covered with greyish-yellow wool; back and waist are
brown. Very long wool on head is very remarkable feature of appearance of barbudo.
On nape and neck the mop of rigid black hair grows as a mane, falling on shoulders
and reaching up to the middle of back and closing neck. At males the magnificent
beard covering throat and partly breast grows (hence the name, literally meaning
“bearded”). Hair of the top part of head are short and light, and consequently
monkey looks as though grown bald though actually only the small part of skin
on head is lack of wool. When monkeys move through grass and bush, long hair
of mane protect body against pricks and scratches. Mane and beard serve for
demonstration in male duels – despite of frightening appearance, males of barbudo
do not enter fight with each other, and confine themselves only to demonstration
of menace and loud cry. In heat of the conflict male can seize by paws a trunk
of small tree or bush and start shaking them, but never rushes to another male.
The face of barbudo is covered with short white wool. Both males and females
have rich eyebrowes – it is a protection against insects and dust, important
at life on the ground. On the face of barbudo there is rather small site of
hairless skin – it is a nose and the bottom part of nose bridge, area between
nose and mouth, and area around of mouth. This skin is brightly coloured, and
the shade of colouring varies depending on physical and emotional condition
of animal. At males the naked skin on muzzle is colored crimson-red color,
at females it is pink. At young animals skin on muzzle is bright pink. At males
there is also a site of naked skin of red color on breast. During menacing
demonstration males can rear on hind legs, showing brightly painted skin of
breast to each other. Thus they raise head, in order not to close breast by
beard, and it gives to their conflict comic similarity to dispute of old men.
In general, by behaviour barbudos resemble African baboons of Holocene epoch,
but differ from them in considerably weaker aggression.
Barbudo has large eyes and good full color stereoscopic sight got from ancestors.
With the help of keen sight animals notice berries and insects making a significant
part of their diet. On hands at these monkeys thick strong nails grow, due
to which animals can rummage in ground in searches of larvae and roots. But
at them the thumb can not oppose to fingers – it is a common feature of New
World monkeys.
Monkeys spend a lot of time in bush and underbrush, therefore sound communication
occupies the significant place in their life. The voice of these monkeys is
rich in various sounds; it includes grumble and grunt, cries of various tonalities
and melodious twittering. Very much advanced hearing helps to warn beforehand
about approach of predators. Having found out a predator, monkeys give the
alarm, squalling shrilly. Their voices serve as the warning for forest inhabitants,
like in Holocene voices of deer, langurs and peacocks notified jungle on presence
of tiger.
In group of barbudo there are some leading males, which copulate with all females
of the group by turns, therefore it is rather inconveniently to define, from
what male cubs are born. Young males had not got to the circle of group “elite”
have no opportunity to couple to females: females do not admit to themselves
males of low rank. It is possible to raise the rank in group by only way: to
protect herd against predators and strangers from other groups, and to concern
loyally to females, occasionally allowing them to steal the found food. Such
behaviour reduces the amount of disputed situations in group.
Once a year the female gives rise to one cub. It strongly differs in colouring
from adult individuals: it has golden wool, and nape and lateral parts of head
are grey. The cub is looked after by the female, but she allows dominant males
to clean wool of the cub and to treat him with found forage when it grows up.
At the age of one year the cub becomes independent, and the difficult period
of its life begins – it becomes “adolescent” to whom children’s pranks are
not allowed. “Adolescents” keep by unite group, and frequently unite with coevals
from other groups, or pass to the next groups one by one, especially if those
groups are small. Three-year old male reaches blossoming of physical force,
and young female at this age already gives rise to the first cub. Life expectancy
of barbudo reaches 15 – 17 years.
This species of mammals is discovered by Momus, the forum member.
Kong (Titanosimia kong)
Order: Primates (Primates)
Family: Para-apes (Parapongidae)
Habitat: Jakarta Coast, South-East Asia, tropical forests.
In late Cenozoic apes were the largest primates. Growth of largest of them,
Pleistocene Gigantopithecus, was estimated at 3 meters. At Madagascar, however,
lemurs of genus Megaladapis approximately equal to gorilla in size lived. In
human epoch large species of primates had suffered serious damage from economic
activity of people and direct chasing. Giant lemurs, undoubtedly, had died
out in early historical epoch because of hunting for them. Gorillas and other
apes became rarity because of hunting and destruction of habitats. Their populations
kept to the end of human epoch in some reserves, appeared too small in order
to restore these species. No one species of apes had lived up to Neocene. But
in Southern Asia evolution prepared the replacement to extinct large primates.
The descendant of one macaque (Macaca) species had passed to ground habit of
life and began the settling in all suitable habitats in tropical zone of Southern
and South-Eastern Asia. In tropical forests of South-Eastern Asia and at the
Jacarta Coast peninsula kong, the largest of ever living primates, had evolved.
Weight of adult male of this species exceeds 700 kg, and his growth reaches
4 meters. The female of this species is smaller – her weight is about 450 kg
at growth up to 3 meters.
Kong is an ecological analogue of Pleistocene ground sloths of South America
and chalicotheres of Old World. It is too massive to climb on trees, and is
exclusively ground species. Only young individuals are able to climb on trees.
By constitution kong is similar to gorilla – this primate has very long arms,
and back is inclined. Tail is not present at this species.
Living in twilight of underbrush, these primates are compelled to use very
appreciable color spots to distinguish relatives. Sciatic callouses of kong
are colored bright red and bordered by contrast ring of white wool. Sexual
dimorphism is expressed not only in size, but also in colouring – male has
black wool with brownish shade, and female is reddish-brown. Additionally male
has brightly colored face – white with bluish spots on cheeks. Face skin of
females is grey. The male’s neck is decorated with magnificent white mane contrasting
with the general colouring of body. At females wool on neck is only slightly
lengthened and darker, than wool on the body.
Jaws of kong are short and wide. Canines at females are very short and consequently
don’t prevent to lateral movements of jaws at chewing. At males canines are
larger and more appreciable – they are used in courtship displays and during
an establishment of domination relations. The diet of kong is diverse and includes
mainly vegetable food. These primates easily pull out plants from the ground
and chew their tubers and rhizomes by strong molars. The significant part of
their diet is made of foliage of young trees. In order to reach it, these para-apes
rise on rear legs and bend trunk and branches of trees to themselves. Kong
likes to feed in shallow water of tropical rivers with soft aquatic vegetation.
The food of animal origin in diet of kong is presented mainly by carrion and
the rests of prey of large predators. Kong differs in massive constitution
and sluggishness, and therefore it is very bad hunter. Its prey includes very
large invertebrates and sluggish reptiles – tortoise or snake insufficiently
quick after nourishing dinner may easily get for dinner to this primate. This
species can easily drive away predators from their prey, using huge force and
superiority in strength.
Kong lives the small groups numbering no more than ten individuals. In group
there is a well defined leader – it is large adult male. Also there are subdominants
in group – one of senior sons of group leader, or male came from the side and
showing loyalty to the leader and supporting him. As a rule, dominant male
descends from other family, rather than females of group.
Kong breeds very slowly. Females turn able to give birth to cub at the age
of about five or six years. Pregnancy lasts till about one year, and till next
three years the cub depends on mother in many respects. The wool of newborn
kong is bright rusty and darkens in course of time. It develops slowly enough,
and can independently move on four paws only at the age of five months. The
cub is fed with milk till about one year, and only since the tenth month of
life starts to use vegetative forage for food. During the life the female of
kong can bring up no more than five or six cubs. At this species of primates
twins are frequently born – one case to approximately 10 – 12 births. Obviously,
this is an adaptation permitting to increase rate of reproduction at these
animals. These primates have well advanced parental instinct, and frequently
the female, too old to give birth to its own cubs, helps young females to look
after cubs, and even transports the cub to the back. Adult kong does not have
enemies, and this primate has an opportunity to live up to 50 – 60 years.
Enkidou
(Enkidou sylvaticus)
Order: Primates (Primates)
Family: Guenons (Cercopithecidae)
Habitat: Southern Asia, plain and foothill woods.
Disappearance or significant reduction of areas of tropical rainforests during
the ice age had caused mass extinction of species connected to these biotopes.
Among irrevocably missed animals there were large primates – apes. Gibbons,
chimpansees, gorilla and orangutan have died out, not having left descendants.
But small species of monkeys were in incomparably best position. Bieng ecologically
plastic they have developed life in light forests of glacial epoch, and when
woods again have covered tropical zone of Earth, wood inhabitants have
appeared among them. Among new wood monkeys the special family of “false apes”
including very large species descended from macacas and convergently similar
to gorillas of Holocene epoch is especially remarkable. In some areas other
large primates, descendants
of
langoors
(Presbytis) widely settled in Asia, compete with them.
The enkidou named after beast-like hero of Sumer epos is huge primate similar
to gorilla having short tail. It is the primate of massive constitution moving
by the ground mainly on four paws, but sometimes rising on legs. In such position
this monkey can make some steps, holding something in hands. Growth of the
rising monkey reaches 3 meters. The body of animal is covered with shaggy black
wool; face skin is naked, colored red. Brightly colored face is a mark of recognition
by representatives of this species of neighbours. However the area of buttocks
serves as not less important recognition symbol at these species, as well as
at many species of monkeys. On buttocks of enkidou there are large sciatic
callouses covered with naked lilac-blue skin at males and bluish-grey skin
at females. These formations have also alarm function, by their appearance
it is possible to determine approximately the status of individual in group,
its physiological condition and readiness for pairing at female. Thus, sciatic
callouses of this animal serve as its original “passport”.
Wool of cubs is colored differently, than at adults: they are light grey, almost
white, with pink face. Sciatic callouses at them are covered with wool. Such
distinctions in colouring have the important value: they serve as the deterrent
for aggressive adult individuals, showing the “special status” of cub in group.
For this reason all pranks and theft of meal at adult monkeys are forgiven
to cubs, and the adult member of clan may have strong punishment for it.
At males of enkidou in mouth long canines used for demonstration at an establishment
of hierarchy jut out. During fights between adults they are not applied, but
at defense against predator or at rescue of the cub got to its claws adult
males inflict terrible wounds by canines. At females canines are much shorter.
Molars of enkidou are wide and knobby – the significant part of diet of this
species include with coarse sappy vegetative food.
Because of massive constitution enkidous spend the most part of life on the
ground though they can swarm up trees. Especially frequently cubs do it, finding
in branches fruits inaccessible to massive adult individuals. This species
of monkeys is omnivorous, equally willingly eating both vegetative food and
meat (if it will be possible to find it). Enkidous willingly eat tubers and
sweetish roots of ferns, soft leaves of bushes, fruits of different trees.
The food of animal origin includes insects (especially they like juicy and
fat larvae of beetles), bird eggs and nestlings, frogs and small mammals. Using
the might and large number enkidous can take away catch of different predatory
animals, or pick up rests of meals after them. During the meat sharing the
severe hierarchy is observed: the dominant male with huge canines tries this
dish first, choosing to itself best pieces. He personally endows with meat
of females with which he is coupled, after them males with whom supports friendly
relations more often, and further all cubs irrespective of whose are they.
After that he leaves a carcass for individuals of the lowest rank and does
not care of its sharing any more. Frequently lowest individuals in hierarchy
get only uneatable pieces of skin and cartilages, and sometimes they get only
smell of meat. If someone from animals of the lowest rank tries to steal a
piece of meat out of turn, the true fight accompanied with loud roar and menacing
grimaces can be fastened. At this time any monkey does not try to steal meat
from fear to get “in the heat of the moment” to the prepotent male. It happens
that in an impulse of anger the male inflicts to members of clan traumas and
mutilations by impacts of powerful hands.
The sharing of vegetative food never happen such noisy and strict because usually
it is enough of it for all ones. However dominant males watch closely members
of clan being always ready to take away from them something tasty, that they
have to found and had not time to hide and to have eaten. At the same time
the leader strictly watches that anybody, especially youngsters, did not take
away meal from cubs. Caring about cubs, the dominant male strengthens his social
status in clan - a lot of females will want to be coupled to him and will support
him if there will be any threat of leadership change in clan.
But the dominant male should fulfil such privileges to the full: at an attack
of predators or at conflict with the next clan he acts in the forefront among
defenders, and at times outcome of the conflict depends on his behaviour.
Time of pairing at enkidou is stretched for half-year falling to rainseason.
At the female ready to pairing sciatic callouses increase and become bright
blue. Usually each male occupying high enough position in clan has one female
to which he couples regularly and supports friendly relations during all life.
The female from such steady pair even can abandon clan together with the male
when the readiness for pairing comes, spending some days at some distance from
the basic group of clan.
Pregnancy at these species of monkeys lasts about half-year. The newborn cub
is covered with wool, at it eyes are opened, and it practically at once can
move head and grip for wool of mother. First time mother constantly carries
the cub at breast. Later it gets over to her back. At bi-monthly age cub climbs
down from mother and starts to play with coevals, at three-monthly age it starts
to try food of adult animals. Half year old cub ceases to suck milk and starts
to spend more time on the ground, moving together with adults. At one-year-old
age the wool of the cub starts to darken: it starts to mature and turns to
the “teenager”. At this time it usually receives the first punishment from
adult animals that marks the ending of the carefree childhood. But connection
between young animal and its mother is kept for long years, even when the female
has other cub. At the age of three years the young female is ready to give
rise to first cub.
Life expectancy at enkidou is 40 - 50 years. For this time up to 6 - 7 cubs
can be born at the female.
Jumping
squirrel guenon (Sciuropithecus semivolans)
Order: Primates (Primates)
Family: Guenons (Cercopithecidae)
Habitat: tropical woods of the Central Africa, highest trees.
At the end of Holocene tropical rainforests of equatorial zone of Earth
have had not best time: aridization and the cold of planet climate during the
ice age has sharply reduced their areas, having caused disintegration of earler
uniform zone of woods to the set of small islands. Representatives of fauna
of these natural communities have died out or their number was sharply reduced.
But some species used original strategy of survival which had helped to stay
number of species at former level: they have simply decreased the size and
at the reduced territory of woods the big population which survival is more
stable could live. Elephants and rhinoceroses from islands of Mediterranean
Sea, mammoths of Wrangel Island and Columbian elephants of islands near coast
North America had used this strategy in Pleistocene. And in forests of Africa
guenons (Cercopithecus) have acted so. One of species of these motley monkeys
had turned to real dwarf in its genus that has permitted to it to survive.
And this tactics had appeared so successful, that later this species had evolved
into some affiliated species occupying different ecological niches. So the
genus of squirrel guenons (Sciuropithecus) which representatives practically
do not exceed the common squirrel by size had appeared on Earth. After
increase of the forest area species of this genus were settled widely, but
the inhabiting at different “islands” of forest had resulted in formation of
different species distinguished from each other by behaviour and habit of life.
Squirrel guenons are very “large-headed” in comparison with other species of
monkeys – it is common feature of all small primates including South-American
marmosets (Callitrix) to which squirrel guenons are similar by ecology. Besides
these tiny primates differ from each other by numerous “ornaments” of hair,
color strips and spots serving for specific recognition.
Jumping squirrel guenon is the typical representative of genus. It lives on
highest trees of tropical wood and has the original adaptation for such life.
This monkey is the excellent hopper; at it has big hinder legs with tenacious
long toes. It easily surmounts distance of ten meters and more by jump. For
small creature it is a simple task: the monkey keeps in air due to long hair
growing on sides and shoulders forming original “parachute”. The magnificent
hairy brush on the tip of long tail helps to drive in flight. By it the jumping
squirrel guenon is convergently similar to monkeys of guereza genus (Colobus).
Pushing off from branch by legs, monkey stretches hands in sides and freely
soars in air, supported by ascending air streams. Having reached up to the
planned branch, it bends body, extends legs forward, deflects head back, and
seizes bark by toes. Due to abilities to “fly” it practically has no competitors
among neighbours, having mastered life on highest trees of tropical forest.
The jumping squirrel guenon is colored contrastly: almost all body is covered
with white wool, hairs of “parachute” are white too, and on back there is black
longitudinal strip expanded on shoulders. At males faces are covered with bluish
naked skin and have red “sidewhiskers” and beard, at females skin on the face
is pink and hair “ornaments” are not present. Besides at males of this species
long canines are advanced. Cubs sharply differ in colouring from adults: they
are covered with brown wool brightening later.
These monkeys keep in crones of the highest trees (at height over 30 meters)
in flights of 20 – 30 individuals. In group the main pair is precisely appreciable,
and other pairs implicitly submit to them. In rear guard of the group there
are young not sexual matured individuals and bachelors. They are afraid of
leader, and at the same time regularly challenge his right on leadership.
At these animals the “language” of sound signals is advanced including up to
30 various “words” like squeal, hoot, twitterling and cries.
Jumping squirrel guenons eat soft fruits, seeds with thin covers, both various
insects and spiders finding them in tree crones. They do not compete to wood
rodents specialized in feeding by firm forages. Besides squirrel guenons are
rather aggressive to them, and rodents, as a rule, avoid places where these
primates keep.
The female gives rise to one cub once of times 5 – 6 months. First time it
grips for wool on her breast, later gets over on mother’s back. While at the
cub “child's” colouring is kept, all individuals in flight irrespective of
rank willingly play with it. But later, approximately at four-monthly age,
colouring starts to brighten: the cub matures. It finally leaves mother at
the age of seven - eight months though from time to time it communicates with
her during all life. The grown up young growth keeps aloof of the leader seeing
threat to the leadership in bachelors. To prove the right of domination the
male resorts to menacing demonstrations: he opens a mouth wide, showing sharp
canines, and squeals shrilly. Thus he shows the force, jumping in branches
or shaking branch on which it is sitting. Usually such reminders on leadership
are bloodless and are limited only to demonstrations but if the opponent is
audacious and is not going to submit, demonstration of forces passes in fight,
being accompanied by bites and peces of pulled out wool. When the hierarchy
is indisputable, the group can food peacefully, except for moments when the
individual lowest by rank does not succeed to hide a lucky find from vigilant
look of the clan leader. Social connections in clan become stronger with the
help of grooming. Squirrel guenons as however all monkeys express this way
the loyalty or favour to the neighbour.
As well as all small animals squirrel guenons live not so long though due to
the brain size they can be considered as long-livers in comparison, for example,
with rodents: they live up to 10 – 13 years.
Representatives of closely related species living in forests of Equatorial
Africa are:
Marsh
squirrel guenon (Sciuropithecus paludiphylus). This primate belongs
to the same genus as previous one. It differs from jumping squirrel guenon
in
more
massive
constitution, shorter extremities, absence of hair fringe on sides and also
in other wool coloring. Colouring of this species is much darker: head of
black color, grey back and white stomach. Along sides longitudinal rusty-brown
strips spread, on hands there are “gloves” of red wool. Around of eyes this
monkey has rounded sites of naked white colored skin. It is the frightening
colouring which is making an impression of huge unblinking “eyes” of any
large animal. Cubs have no such false “eyes”, and they do not have sites
of red wool on the body. Tail at monkeys of this species is covered with
black wool with white hairy brush on the tip.
This species of monkeys is larger rather than jumping squirrel guenon: adult
monkey is young cat-sizeed and male is larger than female. Living in marshy
woods marsh squirrel guenon frequently searches for food near to water: it
is able to catch crabs and tadpoles in wood pools. Also these monkeys are willingly
fed with tubers and greenery of marsh plants. The significant part of diet
of this species includes tree fruits have fallen on the ground, therefore monkeys
of this species frequently watch groups of monkeys of other species, gathering
fruits thrown by them.
Decorated
squirrel guenon (Sciuropithecus ornatus). This representative of
the genus differs in very bright colouring: bright-red body, white crest
on head
at
animals
of both sexes, long tail at the male is white, at the female black. The back
side of palms and feet is colored coffee-brown. Face skin is naked and yellow.
Males have small black “barb”. Cubs are colored uniform brown color; only
face skin is yellow, as at adult monkeys.
Decorated squirrel guenon lives on the trees making forest canopy and practically
does not go down to ground. This species even searches for water in leaves
and crowns of epiphytic plants. Food specialization is available at these species:
fruits of trees of family Sapindaceae including inedible for other monkeys
contain more than half of diet of these monkeys. Fruits of some Sapindaceae
are poisonous being unripe or overripe, but features of digestion of monkeys
of this species permit to eat even them without harm for themselves. Due to
it decorated squirrel guenon avoids food competition to other species of monkeys
and also to rodents.
Blue-faced
squirrel guenon (Sciuropithecus cyanofrons). Distinctive features
of this species is naked bright-blue face skin at monkeys of both sexes and
of any
age, and
also cross-striped black-and-white tails (as at ring-tailed lemur or the
coati raccoon). The body is colored gray-brown color; wool on extremities
is lighter, at males on chin and throat there are long hair form “beard”,
and on head there is black “hat” contrasting with it. Cubs have uniform gray-brown
colouring, and they do not have strips on tails.
This species spends a lot of time on the ground, therefore at them the feature
similar with coati and ring-tailed lemur had developed: these monkeys keep
striped tails vertically, therefore any member of group can notice easily neighbours
among roots of trees. Life of these monkeys is closely connected to trees of
fig (Ficus) genus, which fruits frequently ripen on trunks practically at the
ground level. Other features of behaviour at these monkeys are common for representatives
of this genus.
Thick-foreheaded
obda (Obda pachyfrons)
Order: Hoofed lagomorphs (Ungulagomorpha)
Family Huge lagoids (Titanolagidae)
Habitat: woods and plains of Northern Asia and Beringian mountains.
" Obda" is the name of mammoth at Maris, one of Northern Eurasian folks.
This name is quite warrantly given to the enormous herbivorous animal for features
of external similarity to prehistoric giant. Obda is one of largest ground animals
at the Neocenic Earth: withers height of the adult male reaches 2.5 meters,
body length is up to 5 meters, head length is over 70 cm; animal weights more
than 3 tons (female is a little bit lighter). The gigantic size of animal is
the adaptation for inhabiting in climate of Siberia and Beringia characteristic
in cold snowy winters. Constitution of animal is clumsy: massive body, very thick
legs, every leg with three hoof-like claws. Under fingers there is thick elastic
fatty pat (as at elephant’s leg) which softens steps and expands the support
area of legs. Body is covered with wool – in summer rather short brown with yellowish
sites on cheeks and chest, in winter – dense white one. Wool is especially advanced
on sides; up to winter on back the big fat hump grows lasting up to waist. In
winter “fur coat” on throat and chest rich long “beard” grows, and on bottom
part of sides “skirt” of long wool appears warming stomach. In winter young animals
can hide under this “skirt” with wind as in tent.
On head of animals of both sexes there is massive horn outgrowth spreading
along nose bridge from eyes level (where it is expanded and is partially doubled
covering eyes as if cap peak) to the end of muzzle (where it is narrower and
higher). The outgrowth is used for fights at the relations of hierarchy establishment
and also for snow digging in winter - animal digs out snow by lateral movements
of head, thus the outgrowth works as a shovel. By constitution and presence
of horn this animal is more similar to the extinct hairy rhinoceros (Coelodonta).
Lips of the obda are arranged originally: the upper lip is doubled, and its
halves due to numerous muscles are able to move as if mitten fingers independently
from each other. Such device of lips permits this animal to gather even thin
blades of grass and sticks from the ground. In summer obda eats grass, leaves,
young branches of trees and bushes, in winter it feeds by needles and leaves
of evergreen bushes, digging them out from snow. For digestion of forage rich
in cellulose at animal complex multichamber stomach had developed. Unicellular
organisms inhabiting it help huge animal to digest even young bark of coniferous
trees.
In each jaw of animal there are four incisors and three pairs of wide folded
molars.
Obda lives in herds numbering 20 - 30 animals from which not sexual matured
young growth of different years are about half of number. Herd is under authority
of large male having constant “harem” of 5 – 6 females. Other males in herd
stay at the subordinated position and form constant pairs with mature females.
Females from “harem” of the leading male usually dominate over other females.
Pairing occurs in middle of summer, pregnancy lasts about 10 months. During
rut skirmishes between males usually do not occur as there is a hierarchy in
herd. Adult animals only confirm usual relations of domination with the help
of ritualized movements. Only when young single male tries to challenge superiority
of the lawful leader, rather serious fights can take place though they basically
happen without fatal cases being limited only by might demonstration and short
skirmishes.
Female gives rise to one cub once a year. Cub rising occurs at the end of spring
when herds already are on summer pastures. The newborn cub at once tries to
rise, and after some hours it can freely go after herd. It is fed with very
fat milk and grows quickly: at one-year-old age it weighs about 800 kgs. Usually
shortly before the following delivery at the female milk secretion stops and
the grown up cub completely passes to “adult” food. At this time (usually it
occurs in early spring) the part of cubs perishes if winter appears too long.
Cubs become independent late: the previous cub stays with mother till 2 years,
feeding together with it especially in winter. It becomes completely adult
at the age of 5 years.
Obda makes long annual migrations: in spring herds of these animals move to
the north, and in autumn come back in woods where they spend winter. Migration
usually passes lengthways large rivers walleys where “obda roads” having destroyed
forest vegetation are formed in due course. When old “road” becomes inconvenient
or the herd used it disappears, this place quickly grows with birch or aspen
forest which is later replaced by coniferous trees. Sometimes during migration
separate small herds or family groups unite that interferes with inbreeding.
In spring herds move to estuaries of northern rivers. Tundra during Neocene
has remained as relict zone of separate marshlands covered with low bushes
and in many places taiga approaches almost to Polar Ocean coast. Here obdas
find plentiful forage. In coastal woods obdas eat grass and bushes. Sometimes
they come to sea shallow waters to have a drink salt water (salt is vital to
these animals) or to food with seaweed. At grass feeding obda sometimes kneels
forward legs – at this animal neck is rather short and it is more convenient
to take forage from the ground this way. Sometimes obda foods with river vegetation:
having pulled out from ground tuft of cane or reed, animal “rinses” it in water,
holding in mouth before to to have eaten. In water obdas escape from heat and
blood-sucking insects. Animals swim well and frequently search for forage at
the river bottom digging out by horn tubers and rhizomes of water lilies. Animal
eats plants emerged on water surface.
In autumn obdas migrate to the south, to forests. In conditions of sharply
continental climate when winter frosts fall up to -30°C and northern plains
become covered by one and half meter thick snow layer life in woods looks much
easily. Here they are protected from chill wind and find a lot of forage –
branches of trees and bushes. And at wood edges in silent frosty days obdas
dig out last year's grass and eat the rests of cane sticking up from ice at
the river.
In wool of obda many parasites settle, therefore birds and even bats fed
by them are often and welcome guests in herds of these animals. And in winter
at digged out ground made by obda numerous herbivores – harelopes and rodents
– are feeding.
Life expectancy of obda is great: large males live till 50 years, and the female
- up to 40 – 45 years.
Umingmak (Arctotitanolagus umingmak)
Order: Hoofed lagomorphs (Ungulagomorpha)
Family: Huge lagoids (Titanolagidae)
Habitat: Central and Northern Eurasia, Beringia and Alaska (it is the only
species of hoofed lagomorphs penetrated to New World).
In Holocene fauna of Eurasia herbivorous mammal were presented by hoofed mammals
of various families – mainly by deer and bovines, and also by less numerous
pigs, elephants and perissodactyls (horses and rhinoceroses). As a result of
human activity, hunting and habitat destruction, the majority of families of
hoofed herbivores had disappeared. In Neocene bovines and deer had kept mainly
in southern part of Eurasia and Africa where after human disappearance they
had formed some separate families. To the north of ranges of Central Asia,
in Europe, Siberia and Far East, the fauna of herbivores is presented by various
descendants of pigs, and also by new group of animals – hoofed lagomorphs.
This order of mammals, descended from true lagomorphs, includes animals of
two types – gracefully built harelopes living in savannas, woods, steppes and
mountains, and massive herbivorous huge lagoids to which aurochid from Northern
Eurasia and huge obda from Siberia belong. These animals inhabit woods of Eurasia,
and mountains of Eastern Siberia, Beringia, Alaska and plains of Far North
are inhabited by one more species of massive line of hoofed lagomorphs, the
umingmak.
“Umingmak” is the Eskimo name of musk ox. It indicates a role of this animal
in ecosystems of northern latitudes: it is an ecological analogue of artiodactyls
of Holocene epoch. Umingmak is the herbivorous cow-sized mammal. It lives the
year round in conditions of seasonal sharply continental climate, not making
long migrations like obda. In winter in habitats of this animal there is the
thick snow layer and the temperature sinks up to -30°Ñ, but umingmak is
adapted to survive and to exist normally even in such inclement conditions.
In severe conditions of the North it is better to struggle for survival in
common. Umingmak is gregarious animal keeping in herds of 30 – 40 ones and
sometimes even more. This animal prefers district where wood and plains are
combined. In winter umingmak avoids places where the thick layer of snow falls
– in searches of forage it should dig snow out; therefore these animals go
to forests. Umingmak frequently comes to frozen wood bogs where it is easier
to find high last year's grass.
The animal is well adapted to severe climate of Siberia. Foot of umingmak is
wide: it permits the animal to not fail in snow and to wander freely in snow-covered
woods. Umingmak runs badly – its maximal speed is about 45 kms per hour. But
the predator all the same can not catch it up on friable snow. Back legs of
the umingmak are little bit longer than front ones, and the back is inclined
forward – this animal meets in mountain areas of Central Eurasia and Beringia,
and grazes on slopes. In winter umingmaks often meet in mountains, especially
in areas where the wind blows snow off. They graze stalks of last year's grass
and young branches of bushes. On nose of this animal the cross horn comb is
advanced, assisting to dig snow. It gives to the animal some similarity to
rhinoceros.
At the ground dug by umingmaks other herbivores, snow
harelopes, distant relatives
of the umingmak, feed. Umingmaks frequently search for forage in common with
another large herbivore – Siberian shurga, the huge boar descendant. But they
keep aloof of this aggressive animal, and feed with the rests of grass and
bushes when shurga goes out.
Similarly to their far ancestors, hares, umingmak changes color of wool according
the season. Summer wool of this animal is short and brown, lighter on cheeks
and stomach. On throat and neck of males the “beard” of long hair develops
– it is the secondary sexual attribute characteristic for various species of
huge lagoids. Except for it, males of umingmaks differ in massiver head, rather
than females. In summer wool on back of animal dark “belt” stretches. Winter
wool of umingmak is long and white with grayish shade. At this time at the
animal rich thin underfur develops. In spring shedding umingmaks find coniferous
trees with dried up bottom branches, and scratch against them, leaving on branches
and trunk flocks of shedding winter wool. At this time small birds take wool
of umingmaks to build nests. And larger birds even sit on backs of grazed umingmaks
and pull wool out right at animals. It does not cause them any anxiety: wool
of this animal in general keeps in skin unsteadily, and easily drops out, if
it is pulled out strongly. This feature rescues young umingmaks from small
predators.
Head of umingmak is massive, with thick frontal bones and short strong jaws.
The nasal cavity has grown up, nostrils are able to close – with their help
animal adjusts temperature of inhaled air. Ears of animal are short and rounded
– such feature had developed in cold climate where it is unprofitable to warm-blooded
animal to have strongly extending parts of a body increasing heat emission.
Molars of umingmak are plicated and constantly growing – it is connected to
diet of animal which includes rigid grass and branches of bushes. Forward pair
of top incisors is wide, and all incisors of animal grow constantly. The feeding
animal can bite across small branches of bushes. Umingmak is the exclusive
vegetarian. In summer it eats leaves and grass, occasionally browses branches
of bushes. In winter when it is very difficult to search for food, umingmak
becomes less legible in forage: by wide feet and muzzle it digs out snow in
searches of last year's grass and evergreen bushes. If the layer of snow is
too thick, umingmak can rear and eat branches of trees, basing forward legs
on tree trunk. It can eat even conifer needles.
The simple stomach of umingmak had actually turned to the fermenting chamber
in which symbiotic bacteria live. Umingmak does not chew cud like artiodactyls,
and digestion occurs, basically, in long bowels. For digestion of rough vegetative
forage at the umingmak the volumetric caecum also occupied by bacteria is developed.
Fermented food gets first to caecum, and then to bowels.
The rut of umingmaks takes place at the end of summer when animals are in the
best physical condition. The female ready to pairing emits the special smell
involving males. Around of such female some males gather and persistently follow
behind her. Males establish hierarchy among themselves: they fight, pushing
the contender away from the female by sides and shoulders. Occasionally, when
there are two males of about equal force, they can even bite each other. The
female emitting the specific smell can be attractive to males till some days,
but it is ready to fertilisation till some hours per year. At this time one
of strongest males from her “retinue” simply drives contenders off and protects
the female, frequently pairing with it.
Pregnancy at umingmaks lasts about 10 months, and in spring at the female one
cub (twins are extremely rare, and usually one of twins does not survive) is
born. It is well advanced, and right after birth rises on legs. Umingmaks walk
slowly, and run only in extreme cases, therefore the cub does not lag behind
herd already at the first day of life. It is fed with fat milk and quickly
gathers weight: at the autumn young umingmak weighs already about 300 kgs.
The significant part of cubs had not reached age of two years perishes of winter
fodder shortage. Having survived in first two years, young umingmak has an
every prospect to reach maturity. Males mature at the age of four years, and
the female do it at three years.
Snow harelope (Heterolopa niveophila)
Order: Hoofed lagomorphs (Ungulagomorpha)
Family: Harelopes (Lagolopidae)
Habitat: Siberia, woods and foothills.
The vigorous activity of the person and ice age – these factors both had resulted
to sharp decreasing of the number of hoofed mammals at the territory of Northern
Hemisphere; and they had vanished absolutely at the most part of Eurasia. But
this event had been a stimulus for the further evolution of other groups of
herbivores had been earlier only insignificant addition on the background of
variety of hoofed mammals. Eurasia had became a place of formation of new group
of mammals – hoofed lagomorphs, descendants of ancient and conservative order
including true hares, rabbits and their relatives. At absence of competition
these animals had started to evolve actively, and in Neocene in Eurasia and
North Africa their representatives are very characteristic for fauna of woods
and plains.
Despite of general warming, for Eurasia the severe continental climate with
sharp annual temperature drop is still characteristic. It occurs because of
remoteness of the most part of Siberia from influence of oceans. In summer
in Siberia heat dominates – the temperature rises up to +35°Ñ. But in winter
frosts reach -40°Ñ. Increase of ocean level has resulted in general increase
of amount of deposits. Therefore winters in Siberia became very snow, and in
spring the high water turns to act of nature for local animals. In such conditions
only the animal well adapted to extreme conditions of existence can survive.
One of such inhabitants of Siberia is snow harelope. It is the close relative
of forest harelope, but this species differs from it in larger sizes and heavy-build
constitution. The withers height of this animal reaches 1.7 meters, and weight
is about 300 kgs. Also at the snow harelope there are shorter and rounded ears
covered with fur from outside.
Ancestors of all species of harelopes, hares from Eurasia, differed in ability
to change colouring of fur to winter. Species of harelopes from Europe and
Northern Africa had lost this ability, but snow harelope had kept it. Winter
fur of this animal is rich and snow-white with plentiful thin underfur. Summer
colouring of this animal is brown with light vertical strips on groats and
hips. In summer wool on the head of animal there are white marks on cheeks.
Ears have black tips; in winter fur tips of ears become grey.
On legs of this harelope there are thin fingers covered with the common skin
cover which ends by the common cornificate sole. To winter on legs of animal
“ski” of wool develop, allowing loping on snow. Due to them snow harelope can
move down on slopes of mountains on rear legs bent under.
Snow harelopes escape from predators in flight – they can accelerate momentum
up to 60 kms per hour on firm ground and up to 40 kms per hoour on snow. If
necessary they can defence against predator by impacts of forward legs and
bites. An alarm signal at this species is long shrill whistle.
At snow harelope the precise sexual dimorphism is expressed – male weighs twice
more than female (hence the name including concept “hetero-”- “different”).
At it the “beard” of wool develops: it is an attribute of its good physical
form, and females more willingly pair with the most “bearded” male. This species
keeps in harems numbering one male and some females (usually from three up
to five animals). It is vital necessity – the male is stronger, rather than
females, and in winter it digs out snow to reach last year's grass and evergreen
bushes. Females feed on plants dug out by male. If it perishes from predators,
females are doomed for starvation, but they can join other harems.
In the beginning of winter male pairs with females of its harem. At this time
he is compelled to protect the females from single males challenging to him.
At this time males of snow harelope utter cry similar to low, showing the claim
on females. They fight with each other, kicking and striking impacts by forward
legs. Usually females do not interfere with a course of fight, but occasionally
the strongest female can unite the male, and they drive off the applicant for
a harem together.
At the end of spring at the female two well advanced fawns are born. They rise
on legs in half an hour after birth and at the end of the first day of life
already can run quickly. Fawns differ from adult animals in more contrast striped
colouring and absence of spots on cheeks. They reach the size of adult animals
at the third year of life.
Black-headed
dwarf harelope (Lepolopella nigriceps)
Order: Hoofed lagomorphs (Ungulagomorpha)
Family: Harelopes (Lagolopidae)
Habitat: steppes of Central Asia (up to eastern coast of Fourseas) except for
bush thickets.
Harelopes have replaced in Neocene numerous hoofed mammals became mass species
of herbivorous animals. Having developed plains, forests and woodlands, they
were evolved to set of species with various appearance distinguished by different
requirements in environment. In steppes adjoining to eastern coast of Fourseas
black-headed dwar harelope, one of tiniest representatives of this group of
animals, lives.
Growth of the dwarf harelope at a shoulder is about half meter. This is long-legged,
fragile, gracefully build animal. Legs of this harelope end with sharp hoof-like
claws providing excellent cohesion with ground at run (escaping from enemies
dwarf harelope can accelerate momentum up to 60 kms per hour at short distances
and also sharply change run direction).
Head of dwarf harelope is rather short and high, with strong chewing muscles:
rigid graminoids made the basis of food of this animal. Living in dry and hot
climate dwarf harelope has long ears penetrated with circuit of blood vessels
which promotes effective cooling in heat.
Short wool of dwarf harelope is colored yellowish-brown with narrow close cross
strips. Such colouring is consequence of special defensive strategy of dwarf
harelope: having run any distance animal makes a long jump sideways and hides
in grass having nestled to ground. Striped colouring reliably masks this harelope
from predators.
Remarkable feature of colouring of this species is head parren – black head
sides (areas around of eyes and cheeks). Nose bridge, nape, chin and throat
are colored white. Such colouring of head helps animals to identify each other
among grass and to support visual contact. Keen sight plays very big role in
life of dwarf harelope: with the help of head movements animals can transmit
signals to each other at big distances. If the grass is high, harelopes can
even rear to support visual contact to the neighbour. At cubs head is colored
the same color as body; only at approach of sexual maturity it gradually darkens.
Dwarf harelopes live in herds of 20 - 30 animals at wide grass plains with
small thickets of bushes. There is no well defined leader in herd: the hierarchy
is established by males only in short breeding season. Usually animals spend
day in shadow of bushes and graze since evening till morning with short breaks
for deep sleeping. And animals fall asleep by turns: a little from their number
stay on the alert all the same.
For winter harelops migrate to the east away from Fourseas coast: during winter
rains wool of animals can become wet that usually results in cold. And the
ill harelope is easy catch for predators.
In winter time rich underfur grows at them.
In the beginning of cool winter at harelopes the pairing season begins. At
this time males become pugnacious and intolerant to each other. Rising they
beat each other by forward legs, striking to opponent impacts by forward part
of wrist. Such impacts do not harm fighting animal but at protection of the
cub against small predator female can strike it direct impacts by hoof-like
claws which can wound the aggressor seriously. However protection against the
enemy with the help of hoof impacts is an extremely rare situation: usually
dwarf harelopes seek safety in flight developing at short distance speed up
to 70 kms per hour.
During courtship season males carefully sniff at all females finding ones ready
to pairing. Such condition lasts at the female only some hours per year, and
here it is important to use it in time. Male protects such female within day,
repeating pairing some times. After the female will lose readiness for pairing
and will cease to accept his carings, he searches for new females.
Pregnancy lasts about three months, then in early spring female gives rise
to twin (sometimes even triplet) cubs. They are born very much advanced, with
opened eyes, and after some hours are able to follow herd. At two-day age cub
is equal in speed to adult animals.
Female feeds cubs with milk about 2 months, but already from week age the young
growth starts to try forage of adult animals and change of diet passes gradually.
At the age of eight months young animals get adult black-and-white colouring
of head. They form independent herds and migrate to the east. At this time
females become capable to pairing, and next spring they give rise to first
litter. Males take part in pairing only one year later.
Life expectancy of black-headed dwarf harelopes is rather short and seldomly
exceeds 10 years.
Australian
marsh rabbit (Aqualagus natans)
Order: Lagomorphs (Lagomorpha)
Family: Hares (Leporidae)
Habitat: rivers and bogs of Eastern Australia.
During all Cenozoic in Australia marsupial mammals dominated. Due to absence
of competition they had occupied practically all ecological niches with one
exception. Because of features of cub bearing process marsupial mammals principally
can not develop water habitats. Only in South America the water opossum (Chironectes
minimus) had appeared but it had developed water habitat not in that degree
as, for example, otter or seal. But activity of people had resulted in occurrence
in Australia of big number of placentary mammal species. In Neocene among descendants
of these animals one species successfully occupied Australian waters had appeared.
The place of water predator in Australian ponds and rivers was shared among
different predatory reptiles. But the place of water herbivorous analogue of
beaver and musk rat was earlier vacant. It was occupied by Australian marsh
rabbit – descendant of rabbits introduced by humans. This small mammal had
kept rather recognizing appearance characteristic for rabbits though it has
got some adaptations to unusual habit of life.
This animal is able to swim perfectly and even to dive for 3 – 4 minutes. Strong
hinder legs are adapted to swimming: feet at them are wide. Usually this rabbit
swims having pressed forepaws to sides and making sharp simultaneous movements
by hinder legs. Ears of marsh rabbit are short and rounded, they can close
lengthways. Fur is short and rigid; it is white on stomach and cheeks and brown
on sides and head. Along back some faltering dark stripes pass. Tail is short,
white from wrong side.
Marsh rabbit eats leaves and tubers of water plants. Usually it foods in thickets
of marsh grasses but in case of necessary can dive to the bottom for tubers
of plants - arrowheads and aponogetons. Having dived marsh rabbit digs out
tubers by forepaws. Its ears thus are folded and pressed to shoulders.
Dwelling of this original rabbit is the hole in steep river bank. The entrance
to the hole is placed above maximal water level in the river and usually is
disguised. Holes of different individuals are united to one system and stretch
along river bank to tens meters. The settlement of marsh rabbits usually appears
in place where there are shallows with thickets of water plants and reeds.
In rabbit holes other animals, usually birds and small lizards, frequently
settle.
Settlements of Australian marsh rabbits include usually no more than one and
half tens of adult animals – food resources of river are more limited than
in places where rabbits lived earlier. Three - four times per year number of
colony considerably grows: at does posterity appears. Usually first time doe
of this species rises posterity being half year old. In litter usually there
are four - five cubs. It is less than at rabbits introduced to Australia by
people, but newborns of water rabbits are larger and also develop much faster.
At fortnight age young rabbit cubs start to investigate world around. They
leave a hole and gradually start to try plants eating by adult animals. Rabbit
cubs very cautiously investigate water and make first attempts to swim. It
is the most dangerous moment in their life – numerous predators – turtles,
snakes and fishes hunt for inexperienced young mammals not able to swim quickly.
And at any age predatory birds are dangerous to marsh rabbits and it is possible
to escape from them having dived or having hided in hole.
These animals feed in shallow waters having hided among leaves of marsh plants
rising above water. Because in this case the field of vision is limited at
marsh rabbits the system of sound dialogue is advanced. Resting rabbits communicate
with the help of short clicks. Having noticed the predatory bird flying by
above thickets rabbit utters shout of alarm – prolonged chirring. Having heard
this signal animals try to dive or hide under driftwood. To signal about the
underwater danger marsh rabbits have other signal – shrill whistles.
Despite of care and collective system of enemy watching hardly probable tenth
part from number of newborn rabbit cubs lives up to adult condition. And life
expectancy of adult rabbit seldomly exceeds three years.
New
Azora tardolagus (Tardolagus novazorae)
Order: Lagomorphs (Lagomorpha)
Family: Hares (Leporidae)
Habitat: woods and bushy thickets of New Azora.
In human epoch the fauna of islands had undergone changes: various species of
flora and fauna which could not appear at these islands by natural way had been
introduced to islands. Occurrence of these species has broken natural balance
of island nature, and ecosystem began to develop another way. New species of
animals, descendants of introduced species, had taken the place in these ecosystems.
People had introduced rabbits to Azores (and to other numerous islands of the
Earth). These animals had survived in epoch of global ecological crisis and evolved
to species being a part of new ecosystems of Neocene. At the volcanic island
New Azora very large species of ground herbivorous animals had appeared, the
descendant of rabbit – New Azora tardolagus. It is the largest species of terrestrial
(not connected to sea) inhabitants of island. Tardolagus represents herbivorous
mammal weighting about 50 kg. By proportions of body this animal resembles various
herbivorous mammals evolved at the continents – fossil chalicotheres (Chalicotherium),
megatheres (Megatherium) and Neocene animals - ndipinotheres
(Ndipinotherium)and groundsloth
rodents (Tardimegamys). Tardolagus has long forepaws and inclined
back. Due to “semi-bipedal” constitution tardolagus can rise on strong hinder
legs with wide feet and gnaw branches of bushes and undersized trees. Standing
on hinder legs, this animal reaches growth of one and half meters. During the
feeding it tightens branches by forepaws on which large hooked claws grow, a
little bit similar to claws of ground sloths. At movement on four legs the animal
bases on external side of hands, turning claws by tips to each other. Because
of massive constitution tardolagus moves walking (its name literally means “the
slow hare”), and only in case of danger can run on small distances. Martillas,
large local predators, represent the main danger to this animal. Tail at tardolagus
is very short, long white colored hairs grow on it.
Wool on back, sides and hips is reddish-brown with dark spots forming faltering
longitudinal lines. Lateral parts of head are white, tip of muzzle is dark. Forepaws
from claws up to half of forearm are colored white.
Tardolagus eats rather soft forage – leaves, mushrooms and fruits of plants.
Its muzzle is extended, and forward incisors are rather weak. The animal willingly
eats dropped fruits of trees of laurel and heaths families, and with the help
of claws digs out roots of ferns and other plants.
This animal meets in places overgrown with bushes and undersized trees – at riverbanks,
in gorges and mountains of island. It avoids the areas overgrown with woods of
tropical type with poor underbrush, where there is small amount of food. Tardolagus
keeps in small groups of about ten adult individuals and young growth. This animal
had strongly changed habit of life in comparison with ancestor: it had lost the
ability to dig holes, and only makes beds in bush, or uses natural shelters to
spend the night.
At danger tardolagus prefers to protect itself actively – it rises on hind legs,
bites and beats the aggressor by long claws of forepaws. Thus it growls and opens
mouth, showing incisors.
Breeding rate at tardolagus is very low in comparison with ancestor. This animal
brings posterity once a year. At female one or two large cubs are born. They
are more advanced, than newborn rabbit cubs, but not so advanced, as posterity
of hares. For birth of posterity female leaves the group and searches for shelter
for itself and posterities. The newborn cub of tardolagus is covered with dark
wool, but blind and can not stand. Till first days the female hides posterity
in shelter – in cave or among bushes. The cub develops quickly: at the second
day it opens eyes and starts to explore world around. At week age the cub already
is able to walk. Grown up cubs leave shelter and follow mother to join the group
of congeners.
Young tardolagus begins able to bring posterity at the age of three years. Life
expectancy of this animal is till 20 years.
Beringian hutmaker (Architectona beringica)
Order Lagomorphs (Lagomorpha)
Family Haymakers (Ochotonidae)
Habitat: Beringia, Northern Asia, Big Kuril Isles; in New World from Alaska
up to Greenland, to the south this species penetrates rather far along mountain
ridges.
During Holocene epoch in “shadow” of hares there were their less appreciable
relatives – haymakers (Ochotona). They are similar to rodents, and have no
features of anatomy characteristic for hares – long back legs, permitting to
develop high speed. But these lagomorphs have curious features of behavior
– they gather plants and make of them hayricks for winter (for what they are
named). In addition some species strengthen hayricks by stones and branches.
During the period of transition from Holocene to Neocene this course of evolution
had been picked up and successfully advanced. Evolution successfully realized
the task of survival of these small mammals in some steps. In glacial steppes
of Central Asia where it is a lot of grass, climate is typically continental
and winter is very cold, haymakers had became larger similarly to all polar
animal. Accordingly, they began to provide a plenty of hay, and to protect
it from wind at them feature of behavior was developed: animals began to throw
large branches atop of hay. Surviving in steppe in small colony is rather difficult,
and therefore, animals began to unite to the “towns” numbering tens of adult
individuals: it helps them to increase efficiency of hay reserve.
But to prepare dry grass for winter yet does not mean to spend winter successfully.
Large herbivores may easily find and to have eaten all hay provided by haymakers.
And in steppes of an ice age herbivores were numerous. It were representatives
of hoofed mammals, and also large rodents, new species have appeared in ice
age. To save forage gathered to the winter, separate animals began to protect
stocks, attacking on herbivores, and driving them away by cries, jumps and
even bites. And then some populations had made one more step: they had started
to use stocks of hay as shelters, gradually eating them up from edges and arranging
an inhabited hole in middle of haystack. Gradually among them animals never
settling in holes and other underground shelters had appeared. This step of
evolution had resulted to changes of “house-building” behavior: “having come
off” from dependence on available holes, rocks and crevices, animals have started
to improve “architectural” behavior. They have started to build inhabited tents
separately and did not mix them with stocks of forage. Inhabited tents of these
animals began to be improved in features of design: at their construction a
plenty of branches of trees and prickly bushes was used. For protection against
enemies animals began weaving of branches dense walls, and the internal part
of inhabited tent is covered with grass for warming. So the inhabited tent
is completely separated from stock of forage in behavior of animal, and the
building behavior evolved irrespective of food- providing one.
The Beringian hutmaker had kept features of haymakers in its appearance: it
is similar to short-legged tailless marmot. Auricles of animal are very small,
and eyes are large: it is an adaptation for life in conditions of polar night.
The body of Beringian hutmaker is covered with rich fur becoming longer to
the winter. The fur is colored red with longitudinal black strip along the
back. Wood populations of hutmakers also have dark speckles on back, and often
strip is faltering. The length of body of this animal is about 40 cm; female
is larger than male.
This species builds shelters as tents of branches, warmed from within with
grass and bird feathers. For construction of dwelling the Beringian hutmaker
chooses the prickliest branches, including coniferous trees ones. Building
their shelter, animals accurately cut them out by strong incisors from the
tree-trunk, and drag to building tent. To not give out their presence, animals
cut branches on bushes from the edge of feeding territory. For an internal
lining of house animals gather soft plants, thin stalks of grasses, wool of
sheded animals, and feathers.
In tent up to 2 meters in height there are some inhabited chambers at two or
three floors. The population of one tent makes approximately 4 – 5 pairs of
adult animals, and their posterity. Near to inhabited tent stocks of food for
winter are located: animals gather some stacks of grass, and surround an inhabited
tent by them – it complicates their search by predators. Beringian hutmakers
start to provide a forage approximately from second half of summer. To dry
up a grass and to not spoil it during a rain, some animals in clan constantly
watch weather: if there are signs of rain, they quickly carry off drying grass
under trees. In summer the majority of animals are occupied with preparation
of stocks of forage – they run to glades, cut the overgrown grass, and drag
bunches of picked plants to the common storehouse.
In winter to keep stocks of a forage is vital for clan, therefore in winter
in stacks of hutmakers adult animals and teenagers often “are on duty” – usually
they are males having subordinated position in clan or young animals not having
families.
The adult females already had giving rise to cubs, stand at top of hierarchy
in clan. Per one year at them it may be 2 litters. Cubs are born well advanced,
with opened eyes and covered with wool. They early become independent: already
at week age cubs actively eat forage of adult animals. The young growth, which
has been born at the end of summer, spends winter with parents, and the posterity
from first packs arranges new settlements independently. They quickly grow,
and can give posterity the next year. Life expectancy of Beringian hutmakers
can reach 10 years.
Young animals frequently “convert” for habitation old stocks of forage, strengthening
walls of stacks by branches, and arranging inside inhabited chambers. Doing
it, they as though repeat evolution of dwelling of the hutmaker - from stock
of forage to constant habitation. Usually young females from growing old clan
do it. Gradually they move to the true specially made tents, and their posterity
occupies their habitation.
Sun flying fox (Posteidolon solaris)
Order: Chiropters (Chiroptera)
Family: Flying foxes (Pteropodidae)
Habitat: Zinj Land, tropical woods.
When till the ecological crisis at the boundary of Holocene and Neocene some
birds had become extinct, including predators and the majority of tropical
groups like parrots, bats had an opportunity to occupy new habitats which were
formed in Neocene. Some species of these mammals began much more active in
day time, and in some cases compete to birds as equals. An example of such
species of chiropters is the sun flying fox, the descendant of palm flying
fox (Eidolon helvum), living at the Eastern-African subcontinent (Zinj Land).
Appearing in conditions of partial isolation, this species had appreciably
changed habit of life, had settled in rainforest, and had partly occupied an
ecological niche of animals of forest canopy.
This is rather large chiroptere: body length is about 30 cm, and wingspan is
almost 1 meter. The animal differs in rather massive and strong constitution,
and weighs about 2 kgs. Wings of sun flying fox are shorter and wider, than
at other representatives of order: it is directly connected with the habit
of life of this animal. Sunny flying fox does not like to fly long, limiting
to short flights from tree to tree. In rich tropical wood skill to fly quickly
is not always expedient, but well clambering animals get appreciable advantage
in survival. More often animals of this species prefer to climb on trees, clinging
by hind legs and hooked claws growing on free thumb of wing. Thumb at this
animal is advanced much better, than at other chiropters: it is thicker and
stronger, and claw on it permits this animal to hang under the branch.
Hanging to branches upside down, the sun flying fox dexterously moves on them
in searches of fruits. By manner of movement it resembles the sloth, but moves
much faster than it (this chiroptere accelerates speed up to 5 km per hour).
When it is necessary, animal can even make small jumps from branch to branch
like a monkey. This animal reluctantly flies, making it only in case of emergency
– flying to other tree in searches of food, or escaping from clambering predators.
In flight sun flying fox feels like not so confidently, as other species of
chiropters: it frequently flaps wings (its flight resembles flight of gallinaceous
birds), and sometimes at all prefers to glide from tree to tree similarly to
flying squirrel. The importance of flight in life of this species is so low,
than even the animal having injured flying membrane, can exist and take part
in breeding normally.
The color of wool at sun flying fox partly justifies its name: fur is darkly
yellow (“honey” shade) or sometimes light brown. On stomach wool is brighter,
than on back. Membranes of wings are covered with thin wool on the top side.
On neck of animal there is an ornament: collar of rusty-red fur, especially
advanced at males. Skin of wings is dark brown, contrasting with color of fur.
The head of the solar flying fox looks similar to head of dog with small ears
and large reddish eyes. It is 0rather wide, because animal eats various vegetative
foods and at it chewing muscles are advanced. On skull of this chiropters even
the low longitudinal crest serving for their attachment is developed. This
chiroptere usually eats fruits of any degree of ripening; from unripe up to
began to rot, but it also supplements a diet with leaves and young sprouts.
Sometimes sun flying fox includes food of animal origin to its diet: eggs and
nestlings of birds, frogs and insects. Animal eats fruits simply hanging on
branch near to them, and catches small animals by tenacious hinder legs. Their
flexibility permits to this chiroptere to bring the caught prey by hinder leg
right to the mouth.
As against flying foxes of other species, this species keeps solitarly. Sometimes
on one fructifying tree it is possible to meet several animals of this species,
but they keep separately, not coming nearer to each other. In morning time
animals of this species like to “sunbathe”: climbing up on light up branches,
they stretch wings, having put their inner side and stomach to beams of rising
sun.
The breeding season takes place till all year. Males ready to pairing, involve
females by special “warbles” sounding like long series of clicks. To meet the
female the male prepares special branch: on thick horizontal branch he makes
some odorous marks, using musk secretions of specific gland on chin. The same
secretions impregnate his fur collar, strengthening the smell of animal. Female
ready to pairing, finds these marks in forest canopy, and waits for the male
near of one of them. To draw his attention, the female utters short calls.
After pairing female leaves male’s territory and he continues marriage appeals.
Once per one year at the female one cub is born, occasionally there are twins.
The female holds posterity in the “cradle” formed by stomach and wings. First
some days of life the cub is so helpless, that the female, compelled to fly,
carries him in mouth. Later the cub becomes covered with wool and begins able
to keep by mother during her flight. Approximately at three-week age cub starts
to study to climb on branches. Posterity stays with mother for a long time:
even having studied to fly, and having reached 70 % of weight of adult animal,
the young animal does not leave mother. Because of these circumstances the
rate of reproduction at sun flying foxes is very slow: for one year the female
can give birth to posterity only once (or two times if the first birth happens
in the beginning of current year and the second one happens at the end of year).
But the posterity is teaching by mother in skills of food searching in tropical
wood, learns terms of fructification of various trees and ways of protection
against enemies. Besides the female actively protects posterity in first months
of life, and its survival rate is rather high.
The idea about existence of this species of animals was supposed by Bhut, the participant of forum.
Obda
bat (Synobda medica)
Order: Chiropters (Chiroptera)
Family Common bats (Vespertilionidae)
Habitat: Northern Asia, Beringia. In summer this mammal keeps near to herds
of obda migrating in winter to
southern areas and falling into hibernation.
Large species of animals is the remarkable phenomenon in life of natural community.
Any species of live creatures are an inhabitancy for numerous species of invertebrates
- parasites (their set is named parasitocenosis), but large animal species
become also an original component of inhabitancy even for small (and sometimes
not only to small) vertebrate animals. Among vertebrates of the Neocaenic Earth
the species completely dependent on other vertebrate has not appeared yet but
some species have entered close symbiotic relations with each other. Herds
of large animal obda migrating at spaces of Western Siberia and Beringian mountains
had become mobile dining room for numerous small vertebrates. Large animals
involve clouds of tiny blood-sucking insects eating by various small birds
and also by bats. And one species of bats became practically constant summer
satellite of huge animals.
The obda bat spends the most part of time near herds of obda eating numerous
blood-sucking flies and midges. Activity in extripation of obda parasites of
this species of chiropters is especially important: obda bat flies to hunting
in the evening and hunts bloodsuckers during all night when these tiresome
insects are especially active. But herds of obda constantly move and to not
lose so attractive source of livelihood obda bat has partly torn with ancestral
habit of life and had exchanged homebody life for the nomadic life: these small
mammals spend day in wool of obda; at the same place they leave grown up cubs
for the period of hunting.
Obda bat is small creature: body length of adult mammal is only 7 – 8 cm; wingspan
is up to 25 cm. The body of this chiroptere is covered with short wool of brownish-red
color; on belly wool is grayish-white. Wing membranes and ears are dark and
not covered with wool. Tail is very short and does not exceed hinder legs in
length.
At obda bat there are few outgrowths on muzzle characteristic for many chiroptere
species which at times give to its relatives grotesque and fantastical appearance.
Unique original feature of this species is nose extended to rather long and
mobile proboscis covered with naked folded skin. With the help of this proboscis
obda bat diversifies the diet, finding among wool of obda ticks and other parasites.
Ears of obda bat are large, rounded and almost equal to length of head including
proboscis by size. The special cross groove and two thin muscles on the external
and internal sides of ear permit this mammal to fold and to open ears.
Obda bat spends all day on the body of huge host animal. Usually these mammals
keep on neck, sides and stomach of obda. When animals gather for evening rest,
bats fly to hunt. Flapping by narrow wings they promptly rush between gigantic
obdas, uttering series of thin echolocating clicks and picking up mosquitoes
and midges gathering to herd from nearest swamps. Shouts of bats are perfectly
audible to obdas, and huge animals presume to themselves to relax – there is
reliable guard at them. The keen hearing of bats allows them to hear the slightest
rustle of steps of predators cautiously creeping to herd. Hearing extraneous
sounds bats are nervous and their anxiety forces huge herbivores to be more
sensitive and cautious. Bats benefit by so original neighbourhood too: they
are almost not attacked by predatory birds and animals.
The obda bat is so adhered to mighty host that even breeds in wool of obda.
In the beginning of summer at the female cubs and less often twins are born.
First days female carries naked and blind cub on itself, leaving it in wool
of the obda later. The young growth keeps basically on sides and back of animal
– there it is little bit more safe place: when animal passes wade the shallow
river or grovels obscenely through bushes bat cubs will not fall down from
body of giant. It is not known, whether the huge obda realizes gratitude in
relation to bats settling in its wool. Despite of the “menagerie” lodged in
wool obda continues to live habitual life. Sometimes animals exhausted by stings
of mosquitoes and midges search for rescue from them in rivers. At this time
young obda bats yet not able to fly can simply sink. Partly small size of obda
bat can be explaned by one feature of behaviour which was generated at this
species for protection against similar accidents. If the cub not able to fly
is in danger (for example, the obda on which it is sitting decides to take
a bath or to wallow in sand), female can seize it by claws of hinder legs and
to fly any time taking it, though the “overage” cub is practically equal to
mother by size. In case of need the cub utters the special sound signal, calling
mother; she seizes it by paws and lifts up in air. But it proceeds not for
long time: cubs quickly grow and study to fly. At monthly age young obda bat
can freely fly and hunts insects equally with parents. The next year young
females give rise to their own cubs.
In autumn when obdas migrate to the southern forests bats can abandon them.
Obda bat spends winter farther at the south, gathering for hibernation in caves
and hollows of large trees. Hibernation at them is very deep; thus body temperature
is strongly reduced. After hibernation pairing in early spring takes place,
and bats migrate to the north finding herds of obda and settling on animals
after approach of steady warm weather. They do not have attachment to any certain
herd, and hardly probable obda bat will spend more than one season with the
same herd of host animals.
Striped singing bat (Musicilio striata)
Order: Chiropters (Chiroptera)
Family: Common bats (Vespertilionidae)
Habitat: subtropics of Europe, southwestern coast of Fourseas.
Chiropters had evolved at the Earth right at the beginning of Cenozoic, appreciably
later, rather than birds. Probably, the nocturnal habit of life had helped
these animals to avoid a competition to birds. Chiropters had achieved significant
success in evolution - in human epoch they made up about a quarter of variety
of mammalian species. Among them the species occupying various ecological niches
had evolved: herbivores, nectarivores, insectivores, predators and even blood-sucking
parasites. Chiropters of some species form colonies which number is measured
by millions of individuals.
Chiropters had rather safely gone through global ecological crisis. Among them
highly specialized tropical species, and some species of small total number
had become extinct. But right after stabilization of environment conditions
chiropters had entered new blossoming. Some Neocene chiropters during the process
of evolution had developed very original features of behaviour.
Some kinds of separate genus of chiropters – singing bats (Musicilio) live
in warm-temperate and subtropical areas of Europe. They belong to the number
of medium-sized representatives of group – wingspan at them does not exceed
30 cm, and more often it is even smaller.
These animals have not only echolocation, but also well advanced system of
sound communications. Voices of males of these mammal differ in originality.
At different species of these chiropters at similar sounding echolocating signals
voices in heard part of sound range strongly differ. These animals are able
to utter various sounds – from single clicks and simple monotonous trills up
to rather complex signals, including a sequence of sounds of varying frequency.
Striped singing bat lives in subtropical forests of Europe. It is one of typical
representatives of genus. At these animals wool is colored black, and on this
background from shoulders along the back two wide white strips stretch. Hairless
membrane of wings is also dark. Male and female at this species differ in colouring.
At males strips on back are wider, than at females, and on throat often there
is a white spot. The female of striped singing bat has two thin faltering strips
on back, and besides it is larger a little, rather than male. It is the most
impressively looking species of the genus, other species are colored less brigher.
Muzzle at all singing bats is short and wide. On the nose high leaf-shaped
outgrowth grows, on each side of which chink-like nostrils extended vertically
open. Mobile edges of nostrils are supplied with special muscles which change
width of nostril and density of its closing. “Playing” by edges of nostrils,
singing bats “sing” at the exhalation. They send echolocating signals by mouth,
as usual bats.
At singing bats there are large oval ears with pointed tips and large hircuses.
At striped singing bat edges of ears are covered from external side with short
white fringe.
Wings of these chiropters are short and have rounded tips. Long tail is half
included in interfemoral membrane. Flight of these chiropters is slow and flitting.
Singing bats are active in twilight and at night. They are mainly omnivorous,
and also eat large flying insects (moths and beetles) and ripe fruits of plants.
Various species have preferences in diet, but the species eating more vegetative
food dominate at the south, and at the north migrating insectivorous species
prevail. But even northern species during the fruit ripening time pass to partly
vegetarian diet.
The courtship season at singing bats comes in summer, and one cub is born at
the female in spring of the next year. At each species of singing bats the
specific song differs from those at close species. Males of striped singing
bat involve females with the sounds similar to twittering. Every male displays
itself, hanging on the branch from below and arranging the “concert”. Sometimes
on one tree some males gather, and they organize the original musical competition.
However, at these animals males not always suffer the presence of competitors
– it happens, strong, but badly singing male simply disperses contenders to
near trees. But it has very insignificant chances of success – during fight
it is compelled to interrupt song, and females search for others males. Every
male tries to sing as long, as it is possible, not interrupting. Calling the
female, male utters short abrupt trills. When the female answers his appeal,
male starts to sing actually courtship song – series of whistling sounds. First
“syllable” of his songs is the longest, and each subsequent one is shorter,
than previous, and its tone is higher. During the song performance male stretches
wings and displays to the female its own size.
In the afternoon singing bats prefer to hide in shelters – usually in hollows
of trees or under bark. They do not form big colonies, and search for society
of congeners only during migrations or in courtship season. Striped singing
bat belongs to migrating species, and in autumn, with approach of time of cold
fogs going from Fourseas, it flies to the south – to mountain valleys of Asia
Minor. These chiropters make flights at night, and hide in various casual shelters,
gathering in groups of some tens individuals in day time. The disturbed animals
are very aggressive. They hiss to the enemy and put painful bites.
Some birds may be enemies of singing bats. During night hunting owls attack
them, and large gospodar woodpecker pecks hollows where these animals hide,
and eats them.
Along the coast of Fourseas, in subtropical areas of Europe with soft climate
other species of singing bats are found:
Common singing bat (Musicilio cantans) is the most widespread species of genus.
It lives at the extensive territory: from woods in Central Europe at the west
up to forest areas in deltas of rivers flowing to Fourseas at the east. Populations
from edges of area make migrations, and inhabitants of southern areas – Caucasian
Peninsula and Balkan – are settled and run to short-term hibernation. This
species of chiropters has dim colouring: male is black with brownish shade
and white spot on throat; female has dark brown wool, and it does not have
spot on throat. The only ornament in colouring of this species is thin strip
of white wool at edges of ears. Ears are rounded; nasal outgrowth is wide and
short. Wingspan makes about 25 cm. The voice of male represents a simple trill
of short abrupt sounds.
Greater singing bat (Musicilio grandis) is the largest species of the genus
– at some individuals wingspan exceeds 35 cm, though usually it is less. Colouring
of wool at this species is very light – white with yellowish shade; on head
there is darker “cap” (at the male it is bright red, at the female brown).
This species lives at the south – in mountain forests of Balkan and Asia Minor,
near borders of salt and hot Mediterranean Lowland. The basis of ration of
this animal is made of fruits of trees and large insects. Courtship call of
the male sounds like sharp abrupt serialized singing.
Caucasian singing bat (Musicilio caucasica) inhabits mountain and coastal forests
of Caucasian Peninsula, and migrates for wintering to mountain valleys of Asia
Minor. This small species of bats (no more than 20 cm in wingspan) utters the
melodious modulating singing similar a few to the voice of small songbirds.
This species of chiropters is especially active in twilight and in dawn time.
The wool is colored grey with dark strip along the back; at males on throat
there is a yellowish spot. This species eats insects, and only in middle of
summer passes to feeding on soft and overripe fruits.
Cat-headed
pipistrelle (Hirtofalco felinocephalus)
Order: Chiropters (Chiroptera)
Family: Predatory bats (Carnonycteridae)
Habitat: tropical woods of Central Africa.
After extinction of majority of predatory birds species representatives of
other groups of vertebrates began to develop their ecological niche. Usually
various birds became flying carnivorous creatures, but some species of mammals
from chiropters order successfully competed to them. Some bats passed from
insects to more sizeable forage: they began to hunt birds and small mammals.
Cat-headed pipistrelle is predatory day time species of chiropters, descendant
of one of African pipistrelle (Pipistrellus) species. Because of injurious
habit of life at this animal the structure of body has considerably changed.
Cat-headed pipistrelle had turned to day time predator, and now it hunts birds
and small animals in trees crones finding them with the help of sight. At this
species eyes are very large and directed forward: sight is binocular. Ears
are long, triangular and peaked. The skull of cat-headed pipistrelle is short,
jaws are rather wide: its head is similar to comical portrait of cat. The mouth
of animal can open widely – it is connected to mode of catch killing: this
chiroptere kills caught animals by sting in head. Its molars because of it
have got the special form: they are prolonged and work as edges of scissors
crushing
backbone and skull of catch (the similar adaptation was at fossil “marsupial
lion” Tylacoleo). Outgrowths on nose characteristic for the majority of bats
at this species are small and echolocating abilities are reduced – catch can
hear voice of bat and hide. This animal weighs about 300 grammes having wingspan
up to 70 cm. Colouring of wool is reddish-brown, ears are black and above eyes
there are black stains. On chest of male there is yellowish spot.
This species of chiropters hunt in forest among branches, therefore its flight
differs in maneuverability and high speed. Wings are long but rather wide:
bat easily can change direction of movement and fly round branches. Back extremities
of animal are almost free: the flying membrane is attached only to external
edge of hips; short tail has only small skin edging on its basis. Hinder legs
at this bat are very tenacious, the toe and little toe can oppose to others,
why the grip of this animal becomes doubly stronger. On toes hooked claws grow.
Speed of reaction at cat-headed pipistrelle is so fast that this mammal can
easily grip birds by each paw promptly having flown by through their flight.
Mammal eats caught prey somewhere on branch. Pipistrelle plucks caught birds
by hinder leg holding in mouth (on the contrary it flays small mammals by teeth
holding in paw). The animal eats catch with bones easily chews them by powerful
jaws.
Each pipistrelle has hunting area in forest canopy vigilantly protecting from
neighbours. When two pipistrelles meet at the border of territories, they begin
menacing demonstrations which however do not pass to combat: animals are afraid
to injure flying membrane, therefore teeth and claws are out of use. Animals
open wings, flap by them to the contender’s side and loudly squeal at this
moment, widely opening mouth and showing canines. After this bloodless, but
noisy duel both animals mark borders of territory by urine and peacefully miss
each other.
Twice per year in breeding season males “paste up” on borders of territory
odorous “announcements” for females with the help of secretions of musk glands
on throat. Female ready to pairing hangs to branch near to such mark and waits
when male begins to inspect borders of its possession. Sometimes it even rubs
stomach against male’s mark to interrupt its own smell and to not cause aggression
in it. If the male is to her liking female forms pair with him and moves on
his territory. Pairs at cat-headed pipistrelles are formed not only for breeding
season, but also for all time while female look after posterity – two and sometimes
even three cubs. At this time male is compelled to hunt almost constantly.
Cubs are born naked and blind, but quickly open eyes and become wooly. First
time female carries them on itself but later starts to leave them on branch.
Approximately since the second week of life cubs gradually pass from milk to
meat food – in the beginning they eat semidigested meat, then fresh one and
at last at bi-monthly age they study to catch small animals independently.
Sexual maturity comes at the age of half-year. Life expectancy is about 10
years.
Silent flying wolf (Pterolupus silentiosus)
Order: Chiropters (Chiroptera)
Family: False vampires (Megadermatidae)
Habitat: light forests at the south of Meganesia.
After mass extinction evolution can make unexpected moves and then in ecosystems
very original species of live creatures evolve, occupying in ecosystems a role
unusual from the point of view of the human. It has taken place so in Neocene
Meganesia (the continent uniting Australia and New Guinea). Here predatory
medium-sized birds were pressed by very unusual representative of chiropters
which has received the name the flying wolf.
In Holocene in tropics of Old World large chiropters, flying foxes, had widely
settled. They had received the name only for features of shape, because they
were vegetarians. The flying wolf from Meganesia bears the name more deservedly.
Its appearance does not resemble wolf’s one at all, but this chiropteran has
surpassed its ground wingless “prototype” in injurious bents and dexterity
of prey catching. Flying wolf belongs to suborder Microchiroptera and is the
largest species of bats of Neocene. Wingspan of this animal is about 150 cm,
and weight reaches 1 kg. Body of this animal is muscled, on chest the keel
(atypical formation for small chiropters, which is more typical for large flying
foxes) is well advanced. Wings of flying wolf are long and pointed, and flight
is very fast and maneuverable. The thumb on wing is well advanced and mobile;
with its help flying wolf holds catch when eats it, and also clings to bark
of trees and rocks. The wing membrane of this large species of chiropters in
addition is strengthened by collagen fibers which stretch from bones to edge
of wing.
The tail of flying wolf is approximately equal by length to back extremities.
The interthign membrane completely includes it, but covers back legs approximately
up to knee joint. The membrane of wings also covers legs only up to knees.
Shins of this animal are lengthened and muscled, well adapted for catch seizing.
Toes are mobile and armed with hooked claws from which the small animal cannot
escape. On little toe the especially large claw, with which help the animal
kills the seized animals, is advanced.
The wool of animal is colored brown, on throat there is a spot of yellow wool.
Eyes are led round by rings of white wool. Membranes of wings are colored dark
grey.
Flying wolf hunts in day time and in twilight, and has good color sight. Echolocating
abilities at this chiropteran are substantially lost (this animal does not
distinguish small objects), and express only during the flight in twilight.
During day time hunting flying wolf does not use an echolocation at all (hence
its specific name). In connection with this feature the muzzle of flying wolf
is partly lack of skinny outgrowths characteristic for bats. On nose bridge
of this animal the short vertical outgrowth grows, and small skinny platen
surrounds nostrils from outside. Ears are rather short and triangular-shaped
with pointed tips. They are connected to each other in the top part of head,
and their tips are turned in sides.
This species of bats is an active predator, and eats various vertebrates of
small and medium sizes. More often it hunts birds and small ground or climbing
animals – mammals and reptiles. Large eyes of this predator are shifted forward
and provide good binocular sight.
During the hunting flying wolf grasps catch by hinder legs, and sticks claws
deeply into it. The sharp claw on little toe puts to catch deep wounds from
which it bleeds profusely and perishes. Flying wolf especially frequently attacks
various birds – pigeons, small parrots and large passerine birds. The predator
hunts them not hiding: it frightens away birds, and rushes through their flight,
trying to grasp catch by paws. Sometimes it succeeds to seize two birds at
once. This chiropteran attacks small animals from air, and tries to kill by
claws at once. Flying wolf does not like to hunt among rich trees where the
probability to injure the wing membrane is high. However it dexterously seizes
various tree-climbing animals from large branches and tree trunk.
The courtship season at flying wolf comes at the end of rain season. At these
aggressive bats each individual protects the territory from neighbours, and
because of it pairing can become complicated. The male is larger than the female,
but even in this case there are reasons to him to be afraid of her – protecting
the territory, female attacks wings of the male. Therefore, to constrain her
aggression, male presents the female with small gift – killed small animal.
If the female is ready to pairing, she accepts it, and till some days animals
keep together and sleep on one tree. For these days the male is coupled many
times to the female and drives off other males from her. Then the female banishes
him.
In the beginning of the next rain season female gives rise to one large, but
helpless cub. The first weeks of life she flies to hunting with it, but later
she starts to leave it in tree-trunk hollow. Development of the cub proceeds
about four months. At this time it passes from parent’s milk to meat food and
tries to catch small animals (insects and lizards) itself, not being able to
fly at all. At the age of about 15 weeks the cub starts to study to fly, and
in three weeks it leaves mother and leads independent life.
The young flying wolf reaches maturity at the second year of life. Life expectancy
of this chiropteran reaches 30 – 35 years.
Swift-winged whiskered bat (Setostomops hirundipterus)
Order: Chiropters (Chiroptera)
Family: Molossid bats (Molossidae)
Habitat: South America, edges of tropical forests, woodlands.
Among mammals chiropters concede in variety only to rodents. At the boundary
of Holocene and Neocene the part of species of this group had died out because
of destruction of habitats – mainly because of destruction of tropical forests.
But in any case some species were numerous enough, and in epoch of biological
crisis the order remained various all the same.
In Neocene in plains and light forests of South America herds of herbivores
wander – tapirotheres and huge coursing rodents – deermara and giant
paca.
Herds involve plenty of blood-sucking insects – flies, mosquitoes and midges.
And rather large bat hunts for insects – it is an animal having wingspan of
up to 50 cm and weighs of about 200 grammes. All bats are remarkable by more
or less advanced outgrowths on muzzle, and also by ears frequently of strange
shape – these are adaptations for echolocation. At this bat strange appearance
is added with a plenty of rigid long bristles surrounding its muzzle. For this
feature it has received the generic name “whiskered bat”, and the shape of
wings is a reason of specific epithet “swift-winged”.
Wings of this little mammal are pointed; flight is fast and prompt with sharp
turns. Tail of swift-winged whiskered bat is long; the interfemoral membrane
includes it completely, but hind legs from knee and below it are free of membrane.
This bat eats large insects which gather around of representatives of megafauna.
For prety catching animals of this species have got the special adaptation:
around of their mouth resilient hair grow, increasing trapping surface. The
similar adaptation is present at basket-mouths and lustrer birds of Apodiformes
order, living in Central and Eastern Asia, and it represents an example of
convergence – at representatives of different classes from germs different
by origin organs similar by function are formed.
Muzzle of swift-winged whiskered bat is very freakish. The nose of animal is
short, and narrow nostrils are extended vertically and almost parallel to each
other; the top of nose is slightly below forehead. Edges of nostrils are mobile;
nostrils can open or close as narrow slot. Small eyes are directed forward
and are located at height of half of length of nostrils. Upper lip is splitted
by vertical cut, and its halves can move independently from each other. Edges
of upper lip hang down and are covered on edge with eresilient trapping hairs.
The same hairs grow on the bottom jaw. When animal opens mouth during hunting,
halves of upper lip shift in sides, and the bottom jaw moves downwards. Bristles,
sticking in sides, form a funnel directing the met insects right to the mouth.
During the feeding this bat utters echolocating signals with the help of nostrils.
Ears of this species are small. They are pressed to the head and are opened
as two slits closed on top of the head – tips of ears are densely pressed to
each other, and the forward line of ear adheres to head with the help of skin
membrane.
Body and head of animal are covered with bright rusty wool. Male differs from
female in larger size.
Swift-winged whiskered bat is crepuscular and nocturnal animal not competing
to diurnal insectivorous birds. In day time this bat hides in hollows. It forms
small congestions numbering up to 20 – 30 individuals. These groups have rather
constant structure – no more than 20 % of individuals changes for one year.
Seasonal prevalence in breeding of whiskered bat is not present. One cub is
born by female in any season, but more often it happens in autumn months, in
rain season. Young individuals become capable to breeding at the second year
of life. Up to this time their wool differs by colouring from wool of adult
individuals – it is darker and dimmer.
Megaardvark (Megaardwark armatus)
Order: Aardvarks (Tubulidentata)
Family: Aardvarks (Orycteropodidae)
Habitat: Zinj Land, light forests and tropical woods.
Having African origin, the order of aardvarks never differed in variety, and
distribution of these animals was always limited to area of warm climate. These
animals are specialized in feeding on insects though by biochemical parameters
they are close to ungulates.
When the East-African subcontinent, or Zinj Land had splited off from Africa,
among its inhabitants there were aardvarks (Orycteropus afer). These animals
could go through the time of biological crisis, because they lived not in woods,
but in savanna, and social insects which these animals ate, were kept in enough.
Gradually the climate changed, and the significant part of the area of Zinj
Land was occupied by tropical woods. The descendant of aardvark – the megaardvark,
very large insectivorous animal, inhabits these areas.
Megaardvark is a plantigrade animal moving on hind legs and resembling old-fashioned
reconstruction of any dinosaur. The body length of megaardvark including tail
makes up about 3 meters, and height in a waist is up to one and half meters.
The long tail of animal similar to tail of kangaroo, but thicker, counterbalances
forward part of body. When the animal eats to satiety, in tail the stock of
fat is accumulated. Features of locomotion of megaardvark are reflected in
structure of its skeleton. Sacral bone of this animal is large; some last lumbar
vertebrae and first tail vertebrae are fused with it to unite rigid structure.
Hind legs of megaardvark are very strong and brawny. If necessary the animal
is able to run, but speed of its run is only about 20 kms per hour. The massive
animal can not run long, and runs keeping such speed only about from hundred
to two hundred meters.
If the megaardvark is attacked by predator, the animal prefers to not seek
safety in flight and to protect itself with the help of its main weapon. Forepaws
of this animal are armed with constantly growing thick claws. At this one 3-rd
and 4-th fingers on which claws reach the length of 20 cm are especially strongly
advanced. Usually megaardvark walks, having pressed forepaws to chest. Occasionally
animal bases by claws on the ground, especially during the feeding. But the
main function of claws of megaardvark is not defense, but destruction of nests
of social insects, ants and termites. Besides megaardvark breaks by claws rotten
trunks of trees in searches of grubs, and digs out wood litter. In addition
to insects it eats spiders and even small vertebrates – frogs, lizards and
nestlings of birds nesting on the ground. The megaardvark is convergently similar
to huge stegoechidna from tropical forests of Meganesia, but does not eat the
vegetative forage.
Megaardvark is specialized to eating of insects and other small animals. It
has weak bottom jaw and very small mouth on the tip of long narrow snout. Teeth
are strongly reduced and presented only by three or four pairs in each jaw.
They represent cylindrical formations with thin layer of dentine and easily
erased soft pulp. The animal frays caught animals by them. Tongue of megaardvark
has very remarkable structure. It is very long – the tongue is able to extend
from the mouth more, than to half-meter. The basis of tongue is shifted far
back, and attached to brest, as at anteaters of Holocene epoch. Tongue consists
of elastic muscles, and inside it the thin sinew retracting the tongue in mouth
is stretched. The surface of tongue is covered by large knobs and small corneous
thorns between which glands emitting sticky secret are located. Such tongue
allows not only to extract insects, but even to pull small lizards and frogs
out from holes.
Insects are not safe forage. Many kinds of social insects, which are searched
by megaardvark, are armed sharp mandibles, and ants have also poisonous glands.
Poison of some ants eaten by megaardvark, can easily kill cat-sized animal.
For protection against insects nostrils of megaardvark can close with the help
of special ring muscles. Eyes are another place of this animal vulnerable for
attacks of insects. Eyes of megaardvark are small; they are protected from
casual penetration of insects by dense rigid eyelashes. At megaardvark there
is bad sight, but it does not prevent it to live normally. Sharp sight is not
so necessary for life in rich and shady tropical forest. This animal searches
for forage with the help of very keen sense of smell, and the hearing provides
it the additional information about the world around.
The wool of megaardvark is thin and rough, similar to pig bristle. The whole
body of animal is covered with thick and strong wrinkled skin which can not
be penetrated through by mandibles of insects. Besides on back of this sluggish
animal some rows of dermal ossifications (as at fossil ground sloths like Megatherium)
are developed. The predator, risked to attack megaardvark, risks to break teeth
biting to its back. The adult megaardvark practically does not have enemies.
This animal usually eats termites and ants, destroying their nests. This is
one of few animals of Zinj Land which is not afraid of vagrant ants. Except
for them, megaardvark digs out digging insects from wood litter and searches
for larvae in rotten wood. Megaardvark often feeds on carrion and rests of
prey of local predators. It can not tear off and swallow pieces of meat, but
all the same it finds a lot of edible things on the rests of carcass – megaardvark
eats maggots and grubs developing on it. High acidity of gastric juice makes
such diet of animal safe, protecting it from bacterial infections.
The ancestor of megaardvark was well-known for skill of hole digger. Megaardvark
had practically left this skill. For spending the night the animal digs to
itself non-deep hole in wood litter, usually near the fallen tree. These animals
are solitary ones meeting only for pairing. Seasonal prevalence in breeding
of megaardvark is not expressed. Pregnancy lasts till about 17 months. Once
in two years the female gives rise to one large cub. It has soft skin, and
dermal ossifications are poorly advanced. Therefore the cub of megaardvark
is very much vulnerable not only for predators, but even for vagrant ants quietly
eaten by adult animal. However it is born well advanced, and in some hours
after birth is able to follow mother. The female looks at it within approximately
7 months (at this time she pairs once again). The young megaardvark becomes
able to breeding at the age of about five years. Life expectancy of this animal
is more than 60 years.
Forest horned cony (Ceratohyrax prolongocornis)
Order: Hyraxes (Hyracomorpha)
Family: Hyraxes (Hyracidae)
Habitat: Zinj Land, various kinds of plain and mountain forests.
In early Neocene, after extinction of significant part of species of ungulates,
representatives of earlier not numerous and uniform group of hyraxes had got
an opportunity to show their evolutionary potential. Among them representatives
of separate phylogenetic line of running hyraxes had appeared – subfamiliy
Dromohyracinae in Hyracidae family. These animals gradually passed from burrowing
habit of life to existence in plain district. After great split of Africa the
part of species of this group had remained on continent and evolved to giant
species. And the species isolated at the East-African subcontinent, had changed
much less. At Zinj Land representatives of this basal group of hyraxes had
remained.
In forests growing on plains and in foothills one of such species of running
hyraxes lives. Its distinctive feature is a presence of pair of short horn-like
outgrowths on head; because of this feature the animal has received the name
forest horned cony. This is a pig-sized ground mammal: it weighs about 50 kg
at body length up to 70 cm. Legs of animal are longer, rather than at common
hyraxes of Holocene epoch, and forest horned cony is able to run quickly. Hand
and foot are digitigrade. Despite of massive constitution, animal dexterously
jumps over high plank-buttress roots of tropical trees, escaping from chasing
of predators, and is able to change direction of run sharply. By habit of life
it resembles caviomorph rodents of South America like agouti or paca very much.
This animal lacks tail appreciable from outside.
Bones of upper part of skull of forest horned cony are thickened and form a
kind of helmet. On nose bridge of animal two conic horns directed upwards and
forward grow; these structures are covered with dense cornificated hairless
skin. Both males and females have them, and their application is various: at
males horns serve for courtship tournaments and establishment of domination
out of breeding season, and representatives of both genders also use horns
for digging holes and protection against enemies. The attacked horned cony,
not having an opportunity to escape in flight, attacks on predator, and strikes
impacts by head, ramming the aggressor. Small eyes are slightly shifted downwards
and protected by edges of bone “helmet”.
Colouring of wool is dark brown with white spots on sides of head and on stomach.
Male and female do not differ in colouring. The cub does not have white spots.
Forest horned cony lives in family groups consisting of large male, several
females of various ages and their cubs. Shelter for family of animals is the
hole about half meter wide and over ten meters long, directing under ground
to almost three-meter depth. In hole there is well equipped inhabited chamber
covered by vegetative material, some temporary toilet holes where animals leave
dung and urine, masking their presence, and 2 – 3 carefully disguised emergency
exits.
These animals do not have seasonal prevalence in breeding, and in family group
always there are cubs of various ages. Pregnancy lasts till about three months.
The female gives rise to 3 – 4 well advanced cubs born with opened eyes. They
become sexually mature at the age of about two years.
At Zinj Land close species of horned conies live:
Mountain horned cony (Ceratohyrax crassipygus) lives in mountain areas of Zinj
Land, preferring dry bushy districts and thickets of grasses. It is larger,
rather than forest horned cony, because it lives in cooler climate with expressed
daily and seasonal fluctuations of temperatures. At cold snap this species
runs to not deep dormation. In warm season when food is plentiful, on buttocks
of mountain horned cony thick layer of fat is accumulated; it is used in winter
time. Colouring of wool at this animal is lighter, than at forest species –
straw-coloured with grayish shade on back, hips and waist. Horns on skull are
thicker and shorter, than at forest species. With their help animal can pick
out from the ground stones during the hole digging. This species settles in
pairs, but does not avoid presence of congeners.
Dwarf horned cony (Ceratohyrax minimus) inhabits high-mountainous meadows of
Zinj Land. It is a marmot-sized species weighting no more than 6 kgs. It has
short horn-looking outgrowths of skull, and rich wool has dark, almost black
color without marks both at adult individuals, and at cubs. This animal is
able to climb and jump on rocks, searching for grassy plants. In summer dwarf
horned cony also accumulates fat which is depositing on buttocks and hips.
In conditions of high mountains at downturn of temperature it runs into deep
dormation which lasts not less than 3 months. At this time animal uses the
saved up fat stocks.
Sluggish ventrohyrax, “Zinj sloth” (Ventrohyrax bradipus)
Order Damans (Hyracomorpha)
Family Ashkokos, or Climbing Damans (Ascendohyracidae)
Habitat: tropical woods of Zinj Land.
In epochs of climatic changes some types of vegetative communities may completely
disappear, dooming to extinction the significant part of species connected
to them. But at restoration of acceptable conditions of inhabiting settling
of again formed community can proceed by two ways. In one case it is occupied
with descendants of relic species had escaped in few “refuges” where the former
vegetation had kept. In other cases new habitats are accustomed by the species
earlier not dwelt here. It had taken place, for example, at the East African
subcontinent (Zinj Land) where in tropical rainforests the original analogue
of sloth descending from one local group of animals had appeared.
For Zinj Land the special family of climbing damans, not meeting in continental
Africa, is typical. Its characteristic representative is ashkoko (Ashkoko sylvaticus)
- animal similar by constitution to loris - Asian prosimians of Holocene epoch.
But this animal is not so specialized to tree-climbing habit of life, as ventrohyrax
– another animal living in the neighbourhood.
The name “ventrohyrax” means “a stomach of daman” and emphasizes characteristic
position of this animal – upwards a stomach. Ventrohyrax hangs under the branch
similarly to South-American sloths, for what it had received another name –
“Zinj sloth”.
In connection to unusual way of life in anatomy of ventrohyrax there is a number
of interesting features. Paws of animal are adapted to clinging to branches
– they are rather long, and the structure of hand and foot differs in prominent
features. Hand and foot of animal are lengthened (their length is about one
third of general length of the extremity), along their bottom side strong sinews
stretch, which are attached to strong muscles. For attaching of muscles on
paws bones of ventrohyrax crests giving to bones fantastical appearance are
advanced. Fingers of ventrohyrax are very long and tenacious – animal clings
by them to branches, because claws became very small even at damans – far ancestors
of these animals. On forepaw two fingers are kept (only 2-nd and 3-rd, and
other ones, 1-st and 4-th fingers are reduced and do not touch to branch),
on hinder leg all three toes are kept. Fingers and toes are bent like hooks;
skin on palms and feet is naked, covered with cross plicas and callouses. Due
to this adaptation animal easily moves even on prickly branches. At grabbing
of branch muscles automatically contract, and very strong capture is formed:
even the died animal remains hanging on branch, yet will the decomposition
of corpse begins. Muscles of extremities of ventrohyrax have unusual dark red
color: they contain many myoglobin – the substance accumulating oxygen, therefore
they can work any time in “automatic mode”.
Ventrohyrax spends all life among branches, and extremely seldom goes down
to the ground. On the ground this animal is very clumsy: ventrohyrax bases
on back surface of hooked fingers of forepaws and external lateral surface
of foot (ends of toes are turned under the body). The animal walks on the ground
slowly, and as soon as possible tries to find a tree suitable to life to appear
again in world habitual for itself.
The neck of ventrohyrax is very mobile; due to it animal can eat leaves around
of itself, staying at one place. Muzzle of animal is short and wide; molars
are tuberculous with wide masticatory surface. Chewing muscles of animal are
well advanced: in case of necessity ventrohyrax can bite out small branches.
Ventrohyrax had kept features of dental system characteristic for the daman:
in each jaw it has two constantly growing self-sharpening incisors.
Ventrohyrax is covered with shaggy wool – though it lives in tropical climate,
it may easily catch a cold. In equatorial area each day rain may fall, and
only foliage of trees protects ventrohyrax from it. And in wood canopy strong
winds blow frequently. The wool of ventrohyrax is colored brown with darker
separate locks, precisely imitating the pattern of wood bark. The wool of animal
serves as fine masking. Only it and powerful impacts by claws are its unique
protection against predators. Defending itself, animal seizes branch by hinder
legs and defends by both front paws.
Though ventrohyrax spends all life on branches in tropical forest canopy, this
animal does not make risky and fascinating jumps from tree to tree, and in
general it differs in slowness. At such habit of life three-dimensional sight,
characteristic for damans, loses the value, but the wide field of view gets
much more value for life. Therefore eyes of ventrohyrax look more in sides,
than forward.
Ventrohyrax leads rather simple way of life and consequently does not differ
in high mental faculties. Its brain is too small, and brain department of skull
is lengthened and low.
Though the forest canopy is very productive part of tropical wood, here there
is very rigid competition for food resources. In forest canopy a lot of different
mammals, consuming various kinds of forage lives. The close relative of ventrohyrax,
ashkoko, is omnivorous animal, and other climbing animal, sun
flying fox of chiropters order, eats mainly fruits. Ventrohyrax does not compete to them
due to the diet: this animal eats exclusively leaves. Being a sluggish animal,
not needing for long jumps and fast climbing, and having no big expenses of
energy, ventrohyrax eats poorly nutritious and hardly digestive leaves of plants.
In such way it avoids a competition for food with other animals. The stomach
of this animal is complex and consists of several chambers. In the most voluminous
first chamber there is a food fermentation, in which symbiotic bacteria and
protozoans help ventrohyrax. Ventrohyrax easily digests even dense leaves of
figs, rich in rubber.
This species is social; it keeps in tree crones in groups of one male and several
adult females with cubs. The structure of these groups is changeable, and animals
are not connected with strong ties of attachment (except for female and growing
up cubs). Frequently groups of animals, travelling in forest, unite or break
up. Also there are single males joining groups only for pairing.
Two times per one year the female of ventrohyrax gives rise to one cub. It
is well developed, covered with wool, with opened eyes. Color of wool at the
cub is darker than at female. Till first days of life the cub clings to mother
and eats only her milk. Starting approximately from the week age cub of ventrohyrax
starts to try vegetative food. At first the female feeds it with slurry of
belched and semidigested plants. Young animal receives so necessary gastric
microflora. Gradually the cub passes to feed in vegetative forage.
Young ventrohyrax becomes sexual matured approximately at the fourth year of
life. Life expectancy of this animal reaches 30 – 35 years.
Savanna ndipinotherium (Ndipinotherium crassipygus)
Order Damans (Hyracomorpha)
Family Ndipinotheriums (Ndipinotheriidae)
Habitat: Africa to the south and to the east from savannas of Sahara, Arabia;
flat district with alternating sites of savannas and woods.
The ecological crisis connected to human activity had substantially undermined
biological variety of Earth. And the ice age at the boundary of Holocene
and Neocene became additional test for biosphere. As a result the majority
of specialized groups of animals had died out. Large animals, whose populations
in human epoch became too small and isolated to provide a survival and the
further evolution, also had died out. But ones being “in a shadow” in human
epoch – small species of live beings – had got the main “prize” in struggle
for existence. Damans (Hyracomorpha), small order of primitive hoofed mammals
of African origin, became one of such very perspective groups. From small burrowing
or climbing animals of primitive anatomy animals of several types of constitution
had descended: tree-climbing ashkokos similar to lories, running hyracolopas and massive flathorns. Flathorns are not unique group of huge descendants of
damans. Their relatives are members of original family of ndipinotheriums,
large ground animals of Africa and nearest areas of Asia.
“Ndipina” is the name of gorilla in some dialects of Africa in human epoch.
Savanna ndipinotherium farly similar to this primate dyed out long time ago:
it has massive constitution, short back and rather long forward extremities.
It is very large animal – standing on four legs ndipinotherium reaches three
meters at a shoulder, and about four meters length. Rising on hinder legs,
savanna ndipinotherium can reach branches of trees at five-meter height.
By appearance and habit of life this animal is an original analogue of chalicotheres
of the past and contemporary to it huge groundsloth
rodents from Patagonia.
Savanna ndipinotherium is the social animal migrating to long distances in
large herds numbering up to 60 – 100 animals.
The body of savanna ndipinotherium is covered with rough and rather thin wool
– these animals live in warm climate zone. Wool is mainly straw-coloured, and
on back there is the extended dark brown spot. Its shape may vary at different
individuals – from wide longitudinal strip along the back up to “shabrack”
covering crupper and hips. On shoulders of ndipinotherium there are also dark
spots. On throat and chest of animal the white wool grows – it is the attribute
of sexual maturity, and the throat of young animals is dark.
Back legs of all species of ndipinotheriums are plantigrade and very strong.
The foot of daman, the ancestor of these animals, was covered with soft skin.
At ndipinotherium feet cornificate – it is the adaptation to migrating habit
of life. Large herbivorous species can not feed for a long time at the same
place: it strongly exhausts food resources of district. Due to strong muscles
this animal can easily rear, slightly supporting by forelimbs against tree
trunks. Claws of hinder legs are similar to elephant hoofs, but they are stronger.
On hips of an animal the layer of fat which is spent during fodder shortage
is accumulated. Because of this feature this animal has specific epithet “crassipygus”,
meaning “fat-bottomed”. Tail at ndipinotherium is not present absolutely.
Forepaws at ndipinotheriums combine supporting and grasping functions. Animal
has long hands; the third and fourth fingers are thick, strong and surrounded
with the common skin cover on which bottom side the cornificate skin grows.
At four-legged walking the animal supports on the bottom side of these fingers.
The second and fifth fingers of hand do not reach the ground. They are short,
but mobile – they may partly oppose to third and fourth fingers, operating
similarly to the thumb of human hand. By such paws animal turns in and grasps
branches of trees, which it eats. On forepaws of ndipinotherium small hoofs
similar to nails grow.
Head of ndipinotherium is short, wide and rounded; neck is mobile. Ears of
animal are small. Eyes of ndipinotherium are located on each side of head,
prividing the good circular view. Sight of animal is good: alongside with huge
giraffe ostrich ndipinotherium is one of main “watchmen” in savanna, observing
predators from apart.
By mobile lips ndipinotherium browses soft leaves from branches. But when it
is not enough forage, it can equally easily cut through branches and lignificated
stalks up to 2 – 3 cm thick. It is possible due to the inheritance received
from damans – huge sharp incisors similar to rodent ones. If necessary the
animal defends from enemies by bites: it tries to grasp an attacking predator
by forepaws and to bite through its head or thorax. It is enough one bite to
kill a predator equal by size to lion of Holocene epoch. Due to huge size adult
animal has almost no enemies, and mainly young growth or single animals suffer
from predators.
The rut passes very roughly – males roar, calling females and threatening the
contender. They drive competitors off from chosen females by impacts of shoulders
and heads. When fight becomes especially persistent, forepaws may be used.
Pregnancy at ndipinotheriums lasts about one year, therefore the period of
rut practically coincides the time of cub birth.
In litter there is one cub, but it is very much advanced: it is covered with
wool, opens eyes in some minutes after birth, and in half an hour after birth
can rise on legs and tries to walk. The daily cub does not remain behind adults.
It eats fat milk about half-year, and some months after keeps near to mother.
For this time young ndipinotherium reaches about half of height of adult animal.
When the female becomes pregnant, she drives off from herself the grown up
cub, and young ndipinotheriums enter the most difficult period of life: they
should search for food independently. But, as a rule, in herd there are some
childless females (“nurses”) caring of growing up animals a little.
Ndipinotherium becomes completely adult at the fifth year of life. Life expectancy
of this species in nature reaches 55 – 60 years. In old age the muzzle of animal
appreciably grows grey.
Close species of ndipinotheriums live in Africa:
Coastal ndipinotherium (Ndipinotherium littoralis) lives in narrow strip of
woods at the Atlantic coast of Africa. It differs from savanna neighbour in
small size, gracefuller constitution and long extremities. Growth of animal
rearing on hinder legs, reaches 3 meters. This animal easily rises on hinder
legs and can walk in such position some tens meters. Usually this feature of
behaviour is shown in courtship season.
Muzzle is narrower and extended: animal eats rather soft leaves of bushes and
large grasses growing near water.
Animal is colored grey with dark irregular-shaped spots on sides and back legs.
Throat is white even at newborn animals.
In litter at this species usually it may be two cubs.
Mountain ndipinotherium (Ndipinotherium orobius) inhabits heights of the Central
Africa and its area is isolated from other species of family. It differs in
smaller size, stumpy constitution, rather short extremities and rich wool of
dark brown (almost black) color. It is the smallest species of genus: growth
of adult individual rearing on hinder legs does not exceed 2.5 meters. The
tip of muzzle at adult animals is white, at males it oftenly may be a white
spot on chest.
Animal has massive head and strong jaws: this species eats mainly tough grass
and branches of bushes.
Sea ipopo (Behemohyrax littoralis)
Order Damans (Hyracomorpha)
Family Ndipinotheriums (Ndipinotheriidae)
Habitat: Africa, Zinj Land (ocean coast), coast of Tanganyica passage.
In Holocene sea habitats were populated by mammals of three orders: cetaceans,
carnivors (pinnipeds represent not uniform group, but descendants of several
branches of land predators) and sirens. First two groups were submitted by
carnivorous and planktophagous forms and sirens were sea herbivores. In Cenozoic
it also had exist the order of sea ungulates Desmostylia, but it had died out
long before the people occurrence. Hunting and reduction of places of inhabiting,
caused by human activity, and also climatic changes and “plankton accident”
at the boundary of Holocene and Neocene had resulted in complete disappearance
of the majority of representatives of these orders. Only separate relic species
of “old” orders had remained. But in Neocene mammals had made the second attempt
to develop sea habitats – various groups of mammals in different areas of Earth passed to life in sea independently from each other. So, in Europe the
flattooth, the species of semi-aquatic boars had appeared. Sirens led the origin
from African mammals close to proboscids, and in Neocene the similar situation
had repeated again in Africa. Descendants of damans, also related to proboscids,
are one of succeeding Neocenic mammals. And among them animals leading semi-aquatic
habit of life had appeared.
At coast of Indian Ocean one of representatives of water herbivorous mammals
lives. This large animal is the descendant of damans by anatomy, but by ecology
it represents the analogue of fossil mammal Desmostylus. It browses in mangrove
thickets and at sea shallows, eating leaves of mangrove trees, seaweed and
sea grasses. Systematical position of this animal is between flathorns and
ndipinotheriums, but by features of anatomy this mammal is closer to the last
ones. By size this four-footed animal is larger than rhinoceros, and is externally
a little similar to hippopotamus. Because of it there is its name – sea ipopo
(“ipopo” is the form of word “hippopotamus” distorted in African languages).
Sea ipopos meet in areas of warm climate, and their distribution in many respects
coincides with area of mangrove woods growth. The body of ipopo is covered
with short wool, and under skin the layer of fat improving buoyancy is advanced.
Tail is absolutely not present at this animal. The wool on body is light grey
– it makes the swimming ipopo less appreciable on the background of sea water.
On forehead and nose bridge the black wool grows. At swimming animal they are
above water, and do not give out this animal to underwater predators. The amount
of black wool depends on hormonal level of animal, and it is possible to determine
approximately the status of each individual in group by size of black spot
- at dominants black “cap” is largest.
Legs of ipopo are rather long (near to it the flathorn looks stumpy) but this
animal can not rise on hinder legs (like ndipinotheriums), and moves exclusively
on four extremities. Forward legs are little bit longer than back ones, and
back is sloping. Feet are wide, adapted to walk on fenny dirt and to row at
swimming. Hoofs on toes are flattened, similar to elephant’s ones; under foot
the fat pillow softening steps and increasing the area of support develops.
Due to such structure of legs ipopos easily browse in mangrove thickets where
any predator can not reach. Ipopo swims very well – it lives at all small islands
near coasts of Africa and Zinj Land, and separate animals even regularly reach
Madagascar though a stable population of ipopos had not appeared on this island.
Head of animal is large, rounded, with short jaws. Eyes are located in forward
part of skull. Neck at ipopo is mobile – animal easily feed with branches of
mangrove trees. Ears are rounded, mobile and short. The upper lip of ipopo
is adapted for browsing of parts of plants: it is flexible and represents small
proboscis, as at rhinoceros. The end of lip is slightly forked, permitting
this animal to break even thin stalks dexterously. Simultaneously teeth of
animal enable it to eat food of other kind – containing the big amount of mineral
substances. Incisors in bottom jaw of ipopo are wide, scraping by type. They
have deep roots and constantly accrue. Sea ipopo frequently leaves mangrove
woods and goes to shallows, where eats seaweed, sea grasses and sedentary animals
(in this habit it is the convergention with the European flattooth). Due to
universality these animals can live where mangrove trees do not grow – at the
rocky shallows overgrown with seaweed.
On back of ipopo there is the specific gland – it is the heritage remained
from damans. With its help animals submit each other chemical signals, and
dominants mark territory of herd. In mangrove woods they rub backs against
low thick branches of trees, and among rocks they prefer to mark stones forming
the similarity of eaves under which it is possible to stand.
Sea ipopos wander in herds on coasts, preferring the territories overgrown
with mangrove trees. On rocky sites they also live, but each group has extensive
territory – some times larger, than in mangrove wood. In herd of ipopos strict
matriarchy dominates. Males of this species live one by one or in small groups
in boundary territories of female herds. At bare coasts males live one by one
and follow herds of females.
Pairing at ipopos occurs at any time of year. Pregnancy lasts about 15 months.
In litter there is always one cub. But it is very well advanced, and rises
on legs in first some minutes after birth. It is born on the coast; during
the cub birth the female is surrounded by neighbours protecting her from possible
enemies – at this time they rush even to sea birds. When the cub rises on legs,
the female licks it, and some time does not let neighbours to it. After between
the female and cub connection was established, other neighbours sniff newborn
cub. They as though “accept” it so to the clan, and if necessary protect it
in common. Therefore the survival rate of posterity at ipopo is high. But the
cub depends on mother for a long time – it leaves her only at the third year
of life and then she begins able to bear posterity again. For all life at the
ipopo female it may be 4 – 5 cubs. The young animal becomes able to breed at
the age of 6 – 7 years.
This species has more primitive relative, reed ipopo (Behemohyrax calamophilus) eating riverbank vegetation in rivers of Africa. It lives in marshland at riverbanks.
The basis of food of reed ipopo is made with papyrus and other huge grasses.
This is sluggish peaceful animal little bit smaller by size then its sea relative.
The skin of ipopo has striped colouring – on light brown background there are
vertical dark strips. Due to such colouring animal easily masks in thickets.
Tip of muzzle at animals of both genders is white.
The role of reed ipopo in ecosystems of African rivers is great – these giants,
eating papyrus and other plants, preserve rivers from overgrowing. On lighted
by them sites of thickets of papyrus soon larger sprouts of this plant appear.
Reed ipopo digs out plants from a bottom of river by strong forepaws for what
the claw on internal finger of his forepaw is especially strongly advanced.
By this claw reed ipopo “ploughs” river bottom, and then eats roots, tubers
and sappy stalks of water plants emerged on water surface. As well as the sea
neighbour, this is the social animal. But this animal lives in families in
which there is a male. Male and main female mark territory by musk secretions
of back gland.
Possible, the ancestor of these species lived in rivers and lakes, and then
had settled down the current of rivers at first to deltas, and then to the
sea coast, having the respective alterations in habits and diet.
Fast hyracolopa (Hyracolopa velocipes)
Order: Damans (Hyracomorpha)
Family: Running damans (Dromohyracidae)
Habitat: savannas of Northern Africa.
Unspecialized species of live creatures are one of main reserves of evolution
during mass extinctions. Absence of dependence on any limited resources and
the ability to live in various conditions of environment are main “secret weapons”
of such species. And anatomical primitiveness, characteristic for similar animals,
comprises a potential opportunity for realization of various specializations
in aspect of anatomy.
Among African animals damans (Hyracomorpha) were such group: they had represented
the order of rather ancient and rather primitive hoofed mammals. In Neocene,
after mass extinction, they had formed within the limits of order some groups,
differing by habit of life. One group is presented by massive ground tetrapods
– flathorns from Northern Africa. The second group specialized to tree-climbing
way of life and became analogues of lower primates like lories and lemurs.
It is ashkoko (Ashkoko sylvaticus) from Zinj Land, or the East-African subcontinent.
Except for them, in mountains of Central Africa, savannas of Northern Africa
and mountains of Southern Europe representatives of the third branch of damans
– running antelope-like forms, hyracolopas, live. They occupy an ecological
niche of small and medium-sized antelopes, partly competing with harelopes
of northern origin.
By constitution hyracolopas are similar to typical antelopes, but in closer
examination they have some obviously differences. Tail at hyracolopas is not
present: it was reduced at their ancestors, damans. Rear legs of these animals
are little bit longer than front legs because of what the back of hyracolopas
is slightly inclined forward. This is the heritage of their past way of life:
obviously, hyracolopas evolved in mountains of Central or Eastern Africa, and
such structure of body is typical for animals grazing on hillsides. The structure
of legs uindicates these animals are rock-climbers and runners. Extremities
of damans were plantigrade, and foot of rear leg had formed similarity of sucker.
At hyracolopas such adaptation had disappeared: on back side of rear leg there
is only a callous, permitting to sit on stones. Claws had increased and became
similar to small hoofs. As against horses and antelopes, hyracolopa supports
not only on hoof-like claw, but also on the subungius covered with thick cornificate
skin. Lateral toes became small, and touch the ground at movement only a little.
On forward leg only two fingers from four, characteristic for daman, remained:
second and larger third ones.
Head of hyracolopas is lengthened and low, eye-sockets are shifted in forward
part of skull. Brain cavity is low and long, that clearly indicates on – alas!
– low intellectual qualities of these animals. But at top of skull small sagittal
crest stretches, to which strong chewing muscles attach. In dental system of
hyracolopas features, characteristic for ancestors are kept: gnawing type of
incisors. The bottom jaw is short and a little bit “lengthened” by two large
incisors. With their help hyracolopas can dig out bunches of grass with roots.
If necessary incisors turn to fighting weapon: at the attack of small predators
hyracolopas can bite strongly.
At hyracolopas there are large eyes providing good sight. At mountain species
eyes look forward, rather than in sides: so the three-dimensional sight, which
helps to estimate distance before jump, is provided to an animal. Hearing at
hyracolopas is good; ears are big, rounded and mobile. Sense of smell is advanced
well: nasal cavity is short, but high. Smells play the significant role in
life of these animals: on back of hyracolopas there is the specific gland,
producing an odorous secret. With its help animals mark territory: for this
purpose the hyracolopa rears on back legs, nestles by back to the stone or
tree, and rubs against it.
Hyracolopas live in small herds numbering up to 15 - 30 animals. In herd there
is no strictly expressed hierarchy, and its structure may strongly vary in
due course: the herd may break up, or two herds mix up at one pasture and any
time graze together. For night hyracolopas hide in bushes.
All species of this group of herbivores eat grassy plants, but can eat young
branches of trees and bushes. The stomach of hyracolopas is complex, three-chambered
and similar to stomach of ruminant animals. In forward chamber of stomach the
colony of symbiotic protozoans, similar to those at ruminant animals, develops.
They help animal to digest rough cellulose. The blind gut is poorly advanced.
All hyracolopas have kept a habit, characteristic for damans: animals leave
dung always in the same place. Such behavior has the special role in their
life: by smell of dung animals learn about the congeners, not seeing them at
all. Some animals, living in one group, leave dung in same place.
Fast hyracolopa leads atypical for this genus way of life: it prefers plain
habitats, living in savannas of Northern Africa. This animal reaches height
about 60 cm at a shoulder. Wool of this animal is colored grayish-beige color
with narrow and dense cross strips on body. On all legs there are white “stockings”.
Head of fast hyracolopa is colored brighter, than body: through eyes and forehead
black “band” stretches, which ends wash out on neck; tip of muzzle is white.
Such contrast colouring of head helps to warn congeners of danger: the disturbed
animal highly lifts head, and even rears, looking over district. And colouring
of its head becomes appreciable from afar, serving as an alarm signal for neighbours.
Due to strong rear legs fast hyracolopa can make long jumps, escaping from
the enemy: up to six meters in length and two meters in height. Hoof-like claws
are wide; therefore animal can walk on fenny ground and swims rather good.
Frequently hyracolopas of this species escape from predators in water or in
swamp.
The courtship season at all hyracolopas passes roughly: males gather small
harems of females, jealously preserving them from each other. Threatening the
opponent, they growl and show incisors, having opened mouth widely. If contenders
do not concede each other, they can even fight, biting each other in back and
nape, and striking impacts by forward legs.
Once a year at the female well advanced independent twin cubs are born. They
are able to rise of legs at the first hour of life, and the daily cub does
not lag behind mother. The young growth is colored not so bright, as adults:
cubs don’t have “mask” and white sites on muzzle. They appear only at the age
of about one year. At one and half year age young females become able to breeding.
Males grow longer, and mature only up to age of two years.
Other species of hyracolopas live in mountains, and their area is limited to
one or several ranges:
Rift hyracolopa (Hyracolopa riftica) lives in mountain woods of Central Africa,
where it is the analogue of small antelopes like the duiker. Its growth at
a shoulder reaches approximately 40 cm. Wool of this hyracolopa is colored
dark brown, almost black color, and only tip of muzzle, throat and chest are
white. In mountains where it lives, the weather may be rather cool, and ears
of rift hyracolopa are covered from external side with short wool. This animal
eats various kinds of vegetation: grass, ferns, tubers and bulbs of plants.
It meets in herds numbering up to fifty adult animals.
Wooly-eared hyracolopa (Hyracolopa trichotis) lives even higher, than rift
hyracolopa: on open hillsides with poor vegetation. This animal differs in
dexterity and quickness: it can clamber on almost steep slopes. The wool of
this species is colored brown, and only on throat of the adult male white wool
grows. On back in the area of specific gland long wool grows. For protection
against cold ears of this species are much shorter, than at relative species,
and are covered with long wool from external side. This animal gathers in small
groups and carefully marks borders of territory: fodder resources in mountains
are poor, and the herd should have rather extensive territory.
Atlas hyracolopa (Hyracolopa atlassica) is the species of savannas and foothills
of Atlas Mountains, inhabiting also rocky areas of south-west of Europe. It
partly competes with flatland harelopes, eating rigid plants. The body is colored
yellow, on back there is black longitudinal strip. On rear legs, and at separate
individuals even on front ones, there are black cross strips on white background.
Cheeks and throat are white.
Persian hyracolopa (Hyracolopa persica) inhabits mountains of Near East formed
instead of Persian Gulf as a result of movement of Arabian lithospheric plate.
The area of this species is partly crossed with area of rockloper (Lepotragus
nivalis) from the order of hoofed lagomorphs (Ungulagomorpha). But both species
do not compete with each other because of life in different habitats. Rockloper
prefers dry slopes of mountains, and Persian hyracolopa prefers damper valleys
overgrown with bushes. It frequently lives on hillsides overgrown by wood.
This species lives in herds numbering 10 – 15 individuals. Colouring of Persian
hyracolopa is spotty: general colouring is grey, on sides and groats small
black spots are scattered. Forehead and back part of neck are also black. On
internal side of forward and rear legs there are sites of white wool.
Alpine hyracolopa (Hyracolopa alpina) inhabits mountains of Southern Europe
from Pyrenees to Balkan Mountains along the northern edge of Mediterranean
swamps. It is the largest species of genus, reaching the size of domestic goat.
It differs from other relatives in long wool with rich underfur that permits
this animal to live in high mountains. General colouring of this animal is
grey with black longitudinal strip on back and black spots on groats; legs
are dark.
Forest tapirotherium (Tapirotherium probosciferus)
Order: Odd-toed ungulates (Perissodactyla)
Family: Tapirs (Tapiridae)
Habitat: South America, woods.
Till the evolution process the nature had performed original “experiments”
in various parts of Earth when in different time, on different continents,
but in similar environment live creatures appeared very similar by the anatomy
but descending from completely different ancestors. So, in Cenozoic of South
America representatives of local hoofed mammals, pyrotheres (Pyrotheria), externally
similar to proboscids of Old World and North America had appeared. In Neocene
the situation has partly repeated: till the evolution process pigs in Eurasia
had got a kind of proboscis permitting them to gather food and to utter various
sounds. And in South America evolution of relic perissodactyls tapirs had proceeded.
At them the trunk serving for various purposes also was developed. Proboscid
descendants of tapirs represent the special genus of these animals – tapirotheriums
(Tapirotherium).
The largest representative of genus is forest tapirotherium. This very large
animal distinguishes in characteristic appearance. The animal is a little similar
to the mastodon: the length of body of forest tapirotherium reaches 4 meters
at height at a shoulder over 2 meters and weight up to 3 tons. Back of animal
is slightly curved upwards; head is massive. Legs of tapirotherium are thick,
with wide feet, each of ends by three hoofs. Under toes of the animal the pillow
of elastic gristly tissue impregnated with fat develops. Due to it steps of
tapirotherium are silent. Rather big square of support does not make tapirotherium
a good runner, but permits it to walk on fenny soil and to swim well. Tapirotheriums
live at river banks of northern and central part of South America, sharing
habitats with huge local rodent barocavia (Barocavia). But the competition
between them is softened because tapirotheriums are active in day time and
prefer to feed on land. And barocavias feed on river banks at night. Besides
tapirotherium is able to feed with branches from trees inaccessible to huge
rodent, and even can rear for a short time to do it. Still tapirotheriums meet
in rainforests growing in basin of rivers Amazon and Hyppolite. In rich wood
their residence is easy for determining: here there are wide tracks paved in
wood.
Forest tapirotherium lives in warm climate; therefore it has very thin and
short wool. Animal is colored dark brown color, its head is lighter, and trunk
is cross-striped.
The most remarkable feature of tapirotheriums is the long trunk developed from
short proboscis of tapir. The length of trunk of tapirotherium exceeds length
of skull. Skull of animal is short and high, nasal apertures are shifted back
and upwards, similarly to nostrils on skull of the elephant. Trunk of tapirotherium
is flexible, and animals can tear pick thin stalks of grass or bush leaflets
by it. Also tapirotheriums communicate with the help of trunk: colouring of
trunk is distinct from apart, and animals give to each other signals, swinging
by trunk, lifting or lowering it. Besides with the help of trunk tapirotheriums
utter various sounds: it blows, grumbles and peeps. As against to Holocene
elephants and Neocene pigs of Eurasia, tusks at tapirotheriums are abscent.
Also tapirotheriums distinguish from elephants in short ears. But for large
animal in hot climate there is a big problem of heat emission. For this purpose
at tapirotherium on the bottom part of neck and chest the big plica of skin
develops. In hot weather blood vessels penetrating skin dilate, emitting a
part of heat.
Tapirotheriums live solitarly or in small herds of 5 – 10 animals. Usually
old males keep solitarly at the territory of herd of females with cubs. Young
and mature males out of breeding season form “bachelor” herds.
Pregnancy lasts about 15 months. Female gives rise only to one cub, at birth
of twins one cub, as a rule, does not survive. The cub stays with mother for
a long time: about one year it eats mother’s milk, gradually accustoming to
food of adult animals. After transition to vegetative food young females remain
in herd till all life, and males up to approximately two-year-old age. After
that they leave parental herd and join herd of “bachelor” males. They become
sexual mature at the age of 5 years (at females it happens one year earlier).
Life expectancy of tapirotherium may reach 60 years.
One more species lives in mountain regions of South America: mountain
tapirotherium (Tapirotherium montanicolus). It is smaller species (like usual tapir by size),
adapted to inhabiting in cool mountain climate. At it there are rather short
legs and lengthened trunk: so it is easier to move in cross-country terrain.
Also it does not have plica of skin on neck, characteristic for its large relative,
and it is covered with rich wool. This species is colored brighter than forest
tapirotherium: at it light longitudinal strips on dark brown woolare kept.
It is the display of juvenile colouring. Strips are especially distinct on
shoulders and basis of neck, and around of eyes there are big sites of light
wool on black background. This animal lives in Andes where prefers to live
in mountain valleys. Mountain tapirotherium differs from forest congener in
dense constitution, narrower feet and stronger hoofs. Despite of significant
weight, this animal is able to climb on rather abrupt slopes though it makes
it seldom – usually animals of this species live settled in mountain valleys,
and migrate only at lack of forage or because of inclement weather conditions.
This animal is very numerous near to Totora Lake on Altiplano Plateau in Andes.
Obviously, the close relative of mountain tapirotherium is Chilean
tapirotherium (Tapirotherium chiliensis) – medium-sized animal and rather easy constitution.
Chilean tapirotheriums inhabit narrow strip of land between Andes and Pacific
coast of South America. These animals live in various habitats: from foothills
up to ocean coast. They have long legs and trunk, reach the weight about one
ton and can run quickly. At Chilean tapirotherium there are rather small feet
and hoofs, and also neck is lengthened. It has grey colouring with longitudinal
black strips on groats and back. This species keeps in harems of male (sometimes
two males – main and subordinated) and several females with cubs of various
ages. Populations living near the ocean frequently come to the coast where
animals drink salt water and eat seaweed cast ashore.
Tapiroid
(Tapirocephalus velox)
Order: Even-toed ungulates (Artiodactyla)
Family: Trunk boars (Proboscichoeridae)

Habitat: bush thrickets of Southern Europe.
During global changes in ecosystems unspecialized species able to live in various
conditions and to use for food plenty of forage kinds survive basically. During
change of Holocene by Neocene among few survived hoofed mammals pigs were such
universals and champions in survival. After ice age and extinction of big number
of European hoofed mammals pigs were among animals successfully existed in
narrow zone of vegetation clutched between glacier of kilometer thickness and
heated salty desert of Mediterranean. When glacier had receded pigs among the
first wave of migrants had started to develop free territories and later migrants
from Asia and Africa were compelled to compete with specialized descendants
of wild boar (Sus scrofa).
Boar family had becomed prospering group: among them special family of trunk
boars has appeared at which representatives muzzle had turned to short proboscis.
This adaptation has given appreciable advantages to its owners having expanded
opportunities of food getting. In family even such species started to develop
sea coasts had appeared – it is huge flattooth (Belonoporcus
dolichocephalus).
As against to massive flattooth tapiroid is terrestrial animal. Length of body
of adult animal is about 2 meters, height at a shoulder is 1.7 m. The constitution
indicates fast runner in it: at this animal there are muscled body and long
legs. It can accelerate momentum up to 40 kms per hour at the long distance
and at short time accelerate up to 60 kms per hour. On every leg of tapiroid
there are two large hoofs: characteristic for pigs hoofs on 2-nd and 5-th toes
are underdeveloped and sometimes are absent at all. In appearance of tapiroid
there is a feature making it similar to South-American tapir: massive head
with long proboscis. With the help of flexible proboscis tapiroid can browse
leaves and young branches of bushes containing basic part of its ration. Neck
of tapiroid is rather short but due to proboscis it does not have inconveniences
in daily life. Body of animal is covered with rigid thin wool and on back skin
is very thick – it is its protection against predators. On tips of long ears
there are hairy brushes raising sensory acuity. Eyes at tapiroid are small
and sight is rather weak. But sense of smell at it is very keen: due to presence
of proboscis the olfactory surface of nasal cavity is strongly increased.
Wool of tapiroid is colored brownish-yellow with dark longitudinal strips on
back (such colouring is characteristic for boar piglets). Due to inhabiting
in bush thickets this juvenile attribute became very useful to survival.
In mouth of tapiroid sharp tusks directed forward and in sides grow. At males
they reach significant size. Males of this species apply tusks to duels, establishing
hierarchy or combating for female. During fight opponents simply cling each
other by one tusk and try to tumble down the contender by movement of head
and body pressure. Dangerous wounds this way are completely excluded though
at old males tusks sometimes break. At the predator attack animal uses tusks
as thrust weapon causing to predator deep wounds similar to dabber ones.
Tapiroids live in family groups of 15 - 20 uneven-age animals (young growth
of the current year makes up about half of group) under the leading of large
male. Each group lives in the certain territory on which goes adhering about
constant routes. During movement herd treads and eats young growth of bushes,
therefore in habitat of these animals through continuous bush thickets wide
tracks pass. And the more herd living in certain territory, the wider tracks
left by it are. In tracks graminoids grow basically changing by othergrasses
at the edge of tracks. It gives food to many other bush inhabitants.
Tapiroids eat leaves and young sprouts of bushes, soft grasses and mushrooms.
Because of short neck they can not graze on grass and dig out roots and tubers
as their ancestors, but in bushes they find a lot of suitable forage.
At the watering place animals are cautious: they can reach water only having
bent forward legs, therefore any part of family group stands on the alert while
other animals drink water.
Cubs of tapiroids are born in spring when it is a lot of forage and water.
Female gives rise to two - three cubs covered with short wool. By colouring
young growth is similar to adult animals, colouring is only brighter appreciably
and more contrast. Cub rising occurs almost simultaneously at all females of
family group. Cubs spend first days in shelter among bushes, yet they will
get stronger enough to move after adult animals. On 3-rd - 4-th day they abandon
shelter and herd continues movement.
Females can already rise posterity at two-year-old age, and males though become
mature almost simultaneously with females, receive chance to pairing only at
the age of four years having developed to full might. Life expectancy at this
species is about 25 years.
Nozdrokh
(Probosciaper mammutoides)
Order: Even-toed ungulates (Artiodactyla)
Family: Trunk boars (Proboscichoeridae)

Habitat: woods of Central and Eastern Europe up to Ural.
In Russian annals by the word “nozdrokh” any large animal, most likely mammoth
meant, whose frozen carcasses occasionally had been founded by people, derivating
every possible legends. Therefore the large animal of Neocene epoch, rather
reminiscent by the constitution proboscides missed a long time ago, is named
so.
The extinction of large animals caused by chasing from the side of people,
destruction of places of inhabiting and natural accidents, has released a way
for evolution of groups of animals, not numerous and not variable before it,
having allowed them to realize to the full the evolutionary potential. Among
such “lucky beggars” of Neocene epoch non-ruminant artiodactyl mammals appeared.
In North America dominant large animals peccasons from
peccary family (Tayassuidae) appeared. But in Old World pigs (Suidae) have
reached of the big success: among them ecological analogues of such various
hoofed mammals, as horses, bulls, hippopotamuses, tapirs, and even proboscides
have appeared.
Nozdrokh, the inhabitant of woods of Europe, is a huge descendant of the wild
boar, whose growth at a shoulder reaches 2 meters. It differs in heavy addition
and thick legs. Head of nozdrokh is rather big. Well-known tusks, the main
weapon of boar, at this species had started to increase, resembling tusks of
mammoth: they are bent upwards by half rings. It is convenient to rake by such
tusks a wood litter in searches of roots and acorns. And in winter with the
help of tusks nozdrokh easily scatters snow, reaching up to a last year's grass.
Jaws of animal are rather short and strong, in case of necessity it can feed
even with young branches of trees and bark which it peels out by tusks. As
against to proboscides, at nozdrokhs the “conveyor” of replacing teeth is not
formed.
Snout characteristic for pigs at ancestors of nozdrokh started to grow in length,
and became short, but flexible high-grade trunk. Due to it the animal can lift
even a small acorn from the ground. Inside the trunk there are some pairs of
plicas of epithelium. They serve as original additional “vocal chords”, expanding
a spectrum of sounds uttering by the nozdrokh - from the trumpet roar similar
to elephant’s voice, up to silent chirp and squeak.
Ears of nozdrokh are short – it lives in rather cool climate, and problem of
overheat do not arise at it. The animal is covered with a shaggy rough wool
of brown color. To winter at it the rich underfur grows, permitting to endure
rigorous frosts.
In ecosystem of woods of Europe nozdrokh partly occupies an ecological niche
of large wood hoofed mammals – aurochs and urus. It mainly eats leaves of trees
and bushes, adding to them fern rhizomes, tubers and bulbs of various plants.
In summer nozdrokhs willingly feed near reservoirs, pulling out by tusks whole
bushes of reed. Near reservoirs nozdrokhs spend the hottest days of summer.
Due to wide feet they easily cross bogs, and frequently come to bogs to wallow
in dirt and to cool.
This animal keeps in small herds (up to 5 - 6 animals) under the leading of
adult male. It does not tolerate competitors, and expels young males from herd
when they become independent (at the age of approximately 2 years). Except
for him, in herd there are 2 - 3 sexual matured females and other individuals
are young growth. Young males expelled from family groups, form barchelor groups
of 2 - 4 animals. They more tolerantly concern to each other until will get
own harem. Animals in group constantly communicate with each other with the
help of various sound signals – this way of communication is especially good
in rich wood where animals frequently lose sight of each other.
Female every time gives rise to one cub once per 2 years. It feeds exclusively
with her milk, but in case of danger can expect for protection of any of females
of herd. The cub is born rather weak. It is covered with longitudinal strips
– this is the type of juvenile colouring, characteristic for pigs. First two
days of its life all herd stays at the place, preserving it and expecting while
it will start to walk and will get stronger enough. Gradually strips on its
back become dimmer and disappear to bi-monthly age. Sexual maturity comes at
the age of approximately 20 months, and life expectancy may account about 40
years.
Siberian shurga (Niveaper sibiricus)
Order: Artiodactyls (Artiodactyla)
Family: Trunk boars (Proboscichoeridae)
Habitat: mountains of Eastern Siberia, cold areas of Northwest Asia.
Pigs are one of the most successful animals of Neocene epoch. Due to high ecological
plasticity and using human assistance pigs had considerably expanded an area,
and after human disappearance began to evolve freely, adapting to changing
world and surviving in conditions their ancestors never lived. Some species
had developed woods, others began to
live in bush, third ones became dwarf
inhabitants of ocean islands. Open districts of Eurasia enabled pigs to evolve
to huge species.
In mountains of Eastern Siberia one of such huge descendants of boar (Sus scrofa)
wanders in herds. It is the animal covered with rich wool and named Siberian
shurga (“shurga” is the name of boar at Manchurians). This animal is bigger
than bull – weight of adult males reaches two tons, and females weight up to
1600 kg. Siberian shurga is a relict of congelation epoch kept in conditions
of continental climate of Eurasia. Its ancestors were huge woolly wild boars
(rhinoceros-sized animals lived during the congelation at the border of Holocene
and Neocene). This branch of pigs had early separated from the common branch
of trunk boars family, but Siberian shurga, nevertheless, has well advanced
proboscis.
Shurga is perfectly adapted to conditions of rigorous continental climate.
Winter wool of animal is rich, long and colored gray-brown. It forms something
like the “skirt” protecting stomach of animal from cold wind. Summer wool is
short and dark-brown. On tail of animal long wool grows; that’s why it looks
like horse’s one. Such tail is convenient to drive away blood-sucking insects
breeding plentifully till the Siberian summer. Skin of shurga is thick, especially
on nape and shoulders: it serves for protection against large predators, forming
original “armour”. On back right from nape the big fatty hump stretches up
to waist: it is the stock of nutrients for winter. At males the hump forms
thick fat pillow on head, which is used in courtship demonstrations.
Head of shurga is rather large and massive. Muzzle of animal is flattened from
above; canines stick up forward and are bent upwards. It is an adaptation for
digging of forage from under snow: shurga rakes snow by lateral movements of
head. The snout is extended to mobile proboscis, assisting to gather food from
the ground. The proboscis at shurga was developed independently from representatives
of the basic branch of trunk boars. The proboscis of Siberian shurga is wide
and flat, and its free mobile tip surpasses the length of the top jaw only
a little. It permits to warm inhaled air, and it is very important in habitats
of Siberian shurga for which rigorous winter frosts are characteristic. This
animal tears off leaves of bushes and grass by mobile tip of proboscis.
On cheeks of males thick corneous outgrowths covered with thin wool develop:
they are the attribute of sexual dimorphism and means of protection of males
in courtship tournaments. Eyes of Siberian shurga are small: animal has bad
sight. This animal is near-sighted, but has sharp sense of smell. At Siberian
shurga there are big olfactory chambers inside the trunk, formed by advanced
folded epithelium. Ears are short, covered with rich fur on edges and external
side. Despite of these features, at shurga there is very good hearing.
Hoofs of shurga are wide, under heels of animal fat pillows develop; because
of it the foot of this animal is similar to foot of the elephant. Feet at Siberian
shurga are rather wide; it permits this massive animal to walk on snow and
to not fail. They are covered with thick cornificate skin, therefore this animal
can freely walk on firm stones. Hoofs of III and IV toes are strong. With their
help animal breaks ice crust, wandering on mountains and woods of Siberia.
In winter on feet of animal thick corneous layer accrues, due to which shurga
can walk in snow, not falling.
In summer this animal eats bush branches and high grass. Oftently shurga feeds
on rivers with grass and marsh plants, digging the forage out with the help
of tusks. Shurga has kept habits of omnivorous animal: whenever possible this
animal eats carrion, seaweed and sea animals cast ashore. In winter Siberian
shurga digs snow by tusks, throwing it by lateral movements of head. Animal
can dig out snow up to depth one and half meters. On dugs shurga eats evergreen
vegetation, rests of last year's grass. When thickness of snow reaches several
meters, Siberian shurga migrates to woods. There it is fed on branches – when
snow will wear trees out, their trunks disappear under snow cover and crones
become more accessible. Signs of winter feeding of Siberian shurga are well
appreciable in summer wood: it is characteristically “cut” crones of low trees
bitten in winter up to snow level and lower. They branch plentifully, forming
the umbrella-like crone on low thick trunk.
Siberian shurga lives in forest-tundra and mountains, avoiding extensive marshy
places. In summer this animal frequently visits ponds and rivers, but in the
autumn when the long winter wool starts to grow, shurga moves to dry places.
This animal keeps in herds of 8 – 12 animals which most part females and their
cubs of the first years of life account under the leading of the large male.
Joint search of forage and group protection against predators are an effective
way of survival in rigorous conditions of Siberia. The main enemy of Siberian
shurga is Siberian sabertooth, the large representative of felines, specialized
for large prey hunting. Usually it attacks these animals in summer when it
comes from the south with herds of obda. In winter a lot of animals perishes
from fodder shortage and cold. The young growth suffers especially strongly
– in first winter of life about half of the animals, born the previous spring,
perishes.
The rut at Siberian shurga begins with the first frosts in the beginning of
winter. During the rut males are aggressive. They determine the hierarchy with
the help of tournaments during which they beat each other by tusks to tusks,
having lifted heads. Corneous outgrowths on cheeks protect head from casual
impacts. At this time they strongly puff trunks and roar.
Usually male gahthers a harem of several females, and preserves it against
contenders while females are ready to accept its courtship. Male shows fat
stocks to the female, turning sideways to her and shaking head. Simultaneously
turn by side is an action which extinguishes aggression of the female: the
male demonstrates to her the spot vulnerable for tusk impact.
After the rut males and females unite in herds – it is easier to survive in
winter this way. Pregnancy lasts about seven months (it is rather short term
for so large animals). The female gives rise to two – three cubs weighting
about 80 kg. At once at birth they are covered with wool, active, are able
to run at once (piglets spend first days of life in shelter). In two days cubs
of shurga do not remain behind herd. They grow quickly, being fed with fat
milk. At bi-monthly age cubs start to try vegetative food, and to the end of
summer completely pass to diet of adult animals. To the autumn young animals
grow weight about 400 kg. Due to coordinated group defense against predator
their survival rate is good, but in winter a plenty of young growth perishes
from frost and famine – adult animals are not able to protect them from it.
Usually young animals had reached two-year-old age, have good chances to reach
sexual maturity and to breed. Maturity comes at six years age at males, and
in seven – eight years at females. Life expectancy reaches 50 years.
The idea about existence of this animal was stated by Momus, the forum member.
Porceratella
(Porceratella bovina)
Order: Even-toed ungulates (Artiodactyla)
Family: Horned hogs (Porceratidae)

Habitat: tropical woods and marshlands of Zinj Land.
Mass extinction of the majority of groups of hoofed mammals at the boundary
of Holocene and Neocene became mainly the result of human activity. Only most
unpretentious and ecologically plastic species have managed to survive, having
kept as small populations. After disappearance of mankind they have started
to fill in released ecological niches.
The geography of planet in Neocene epoch has changed, though it has remained
rather recognizeable. From Africa along the “seam” of Great Rift Valley the
large land site had “broken away” – Zinj Land. And here one of species of hoofed
mammals, river hog (or bush pig) (Potamochoerus porcus), had turned to the
interesting creature similar to bulls of Holocene epoch.
The river hog had small bone knobs before eyes. Evolution had picked up and
had developed this “idea”, and the descendant of river hog, porceratella, had
acquired a pair of sharp horns directed forward and a little in sides. It is
interesting, that at the continent descendants of bush pigs either have lost
rudiments of horns completely or have acquired one large unpaired horn similar
to horn of brontotheres. All these animals belong to the separate family of
horned hogs (Porceratidae).
Osseous horns of porceratella are precisely separated from each other, and
resemble horns of bovids. They are filled with friable bone tissue lowering
their weight, but their top layer is enough strong and they could be used as
the tournament weapon, and for protection against predators.
Externally porceratella resembles small cow with back curved upwards. Height
of animal at a shoulder is about one and half meters, weight of the male is
about 500 kg, female up to 400 kg. The wool is short, rather thin, brightly
colored: back is rusty-red, sides are white and the bottom part of body is
black. The area under tail is covered with white wool. Behind on hips between
sites of white and red color there passes a black strip. Wool on throat is
also black. Around of eyes there are small white “glasses” (sometimes they
may be absent), on tips of ears there are rich black hairy brushes. The tail
is covered with rich white wool, its tip is black.
The nose has turned to small mobile proboscis, shorter, than at saiga or tapir.
Characteristic for pigs ossifications in snout are reduced and the animal can
not dig out by it something from the ground, but can pick up edible parts of
plants. Due to proboscis the animal can utter various blares. Also the big
surface of nasal ducts promotes development of sharp sense of smell. Sight
plays smaller role in life of porceratella: the bases of horns form protective
“eyebrows” which prevent it to see object that is directly ahead: to make it,
the animal raises head a little.
On the top jaw there are sharp tusks directed forward and in sides. With their
help the animal digs ground in searches of food. In the case necessity porceratella
can put deep wounds by these tusks.
Hoofs can move apart, due to what porceratella is able to walk along fenny
marsh ground. Besides it does not avoid water: it willingly swims, feeds with
water plants, and searches in water for protection against ground predators.
Porceratella keeps in herds numbering 15 - 20 ones. Over the herd is predominated
by the female, and other members of herd are also females. Males keep in herd
only while depend on mother. The leader female expels become adult males. Males
form barchelor groups of 5 - 6 animals in which the not strict hierarchy is
observed. But in season of pairing (in the middle of rain season) they become
aggressive, and can even kill each other in tournament fights.
The posterity is born in the beginning of dry season, when there is no danger
of flooding of forests. In litter there are usually two cubs, at young females
only one cub is born. At the age of approximately three months at young animal
horns start to grow, and half year old males are already compelled to lead
independent life. They completely develop and become able to breeding at the
age of 2 – 2.5 years.
Bikifaru (Dilophoporcula rhinocerotoides)
Order: Even-toed ungulates (Artiodactyla)
Family: Horned hogs (Porceratidae)
Habitat: Zinj Land, light forests and bush thickets in the north part of subcontinent.
Artiodactyl mammals in Holocene turned to favourite objects of people hunting.
Human activity has resulted to strong reducing of the number of various gregarious
animals (mainly bovids), and the majority of species of these animals had disappeared.
Only few relic species kept in Asia and South Africa. The place of small swift-footed
antelopes in Neocene was occupied by various animals of new groups: harelopes
of lagomorph group and hyracolopas, running descendants of damans. And the
place of larger herbivores – rhinoceroses and buffaloes – representatives of
other groups of mammals, various descendants of damans, had shared. At the
East-African subcontinent named Zinj Land, large descendants of damans are
represented only by one semi-aquatic species. At the subcontinent the large
descendants of pigs occupied ecological niches of ruminant animals had appeared.
One of the most widespread species is bikifaru, large species of horned descendants
of pigs.
This animal is larger relative of porceratella, which leads migrating way of
life and differs in rather graceful constitution. This is stout cow-sized animal,
also similar to cows by proportions. At the short distance bikifaru can accelerate
momentum up to 50 kms per hour, but it can not support such rate for a long
time. Hoofs of this animal can move apart - bikifaru frequently feed on juicy
marsh vegetation at the fenny ground. Also this animal is not afraid of water
and is able to swim well.
Wool of animal, living in hot climate, is rough, short, sandy-yellow colored.
On back and hips there are some dim darker spots. The tail of animal is long
– with its help grazing bikifaru drives off blood-sucking insects. On tail
there is big black-and-white brush of long hair. Using it the animal submits
signals to congeners: the tail with hairy brush lifted upwards, well seen from
apart, is the alarm signal, clear not only for bikifarus, but also to their
neighbours.
Neck is moderately long; head is rather small. But the structure of head of
animal has determined its name. “Kifaru” in suachili means “rhinoceros”, and
“bi …” means “double” in Greek. On head of this animal not horns, as at porceratella,
but two longitudinal bone crests grow. These outgrowths are too thin to be
used for combat, and serve more for demonstration. Also they help to move apart
prickly branches of bushes among which bikifaru lives and feeds, and protect
eyes from damages. Crests are semicircular; on nape they stick back like two
horns. These crests are covered with thick rough skin and short wool, and frequently
grow very large – at old males the size of crests can be almost equal to the
seen sideview of head. Crests of males have a border of black wool on edge,
and due to it are appreciable from apart. At females on crests there are only
separate black spots, and at young animals colouring of crests is the same,
as at all body. The width of black border depends on level of testosterone,
and this feature helps males to estimate force of each other, not entering
combat. Females also prefer to couple to males, at which there is more of black
wool on crests. Also at males at edges of crests knobby osseous outgrowths
develop. At males the small tusks, inflexed and turned in sides, also are advanced.
This is the weapon for demonstration, but the defending animal can strike a
predator by lateral movement of head. At females tusks are smaller and are
visible only when the animal opens mouth.
Lips of bikifaru are mobile, and the muzzle is extended to small proboscis.
Inflating proboscis and contracting nostrils, animal can utter various sounds.
In bushes where visibility can be very bad, this animal communicates with congeners
almost exclusively with the help of sounds.
Bikifaru is strict vegetarian; it eats leaves of bushes and branches of young
trees. Jaws of this animal are rather weak; therefore the animal does not eat
rigid grass. Due to the structure of hoofs bikifaru can graze in bogs, where
it eats leaves of huge grasses and gathers floating plants from surface of
water.
This species lives the herds numbering up to 20 – 30 adult animals. The herd
is ruled by large male which constantly keeps up observance of hierarchy. Between
males there is rather rigid struggle for leadership: the dominant male can
couple to the majority of females of herd. Males standing a rank below the
leader also have opportunity to have posterity, especially in big herd. The
old animals, not able to support by force the place in hierarchy, quickly fall
to the bottom step of hierarchy of herd and only the young animals recently
becoming independent can be lower than they are. Old males usually keep in
back part of herd, and fall victims of predators the first.
Seasonal prevalence in breeding of bikifaru is not present. The female ready
to pairing involves at once many males which arrange combat for the right to
couple with her. Pregnancy lasts about eight months and is finished by birth
of one or two cubs (twins are born in third of cases). The cub is born advanced
and can walk at once. It differs from adults in absence of longitudinal crests
on muzzle. Its colouring is brighter, rather than at adults – on sides of young
animal there are black spots, merging to faltering longitudinal strips. At
growing animal they grow pale and vanish on the most part of body. At the animals
living in richer bushes and light forests in south of area, at the border of
rainforests, spots are kept to all life. These animals, larger than northern
ones, are classified as the subspecies Dilophoporcula rhinocerotoides maculata.
Brontothere
hornhog (Porceras brontotherioides)
Order: Even-toed ungulates (Artiodactyla)
Family: Horned hogs (Porceratidae)

Habitat: savannas and light forests of Southern Africa, dry sites, foothills.
Presence at bush pigs (Potamochoerus porcus) of small bone outgrowths before
eyes looks insignificant attribute at first sight. But evolution had picked
up this attribute, and it has received the development at descendants of these
animals: at them on head large bone outgrowths of various shapes have appeared.
So on life arena the family of non-ruminant artiodactyls, horned hogs (Porceratidae),
has appeared. At the representatives
of this family living at the Eastern African microcontinent, there is the
pair of horns a little bit similar to horns of bovids. And at inhabitants of
continental Africa on the head mainly unpaired outgrowths were developed, making
these animals little bit similar to rhinoceroses extinct at the end of Holocene
and brontotheres dyed out to end of Paleogene. In continental Africa these
animals are spreaded mainly to the south from zone of tropical woods, in damp
savanna and mountains of Southern Africa.
The brontothere hornhog is one of the most typical representatives of the family,
widely widespread in rather dry habitats. By constitution it is similar to
bull (growth at a shoulder is up to one and half meters), but its head strikingly
differs from bull’s one. It is larger, with rudimentary proboscis developed
from characteristic snout of pigs. The most remarkable feature is the presence
of huge osseous horn. It is formed by pulled and grown together pair bone knobs
characteristic for ancestor. The horn rises perpendicularly to forehead; it
is expanded in sides and partly doubled at top. It is rather demonstration,
than fighting weapon. But with the help of strong cervical muscles the brontothere
hornhog can hook by horn the attacking predator and reject it aside. During
intraspecific tournaments males of this species butt, having inclined a head
aside, and resting by bases of horns. But usually they avoid direct fight,
and only show horns to each other. At females horns are shorter, than at males.
Wool of this animal is very short. On chin hair are a little bit lengthened.
The horn is hairless, covered with cornificate skin. The basic colouring of
body is light grey, only on nape lengthened black hair form similarity of mane,
and on long tail there is black hairy brush.
In mouth there are two long incisors similar to teeth of fossil rhinoceros
Chilotherium: animal uses them to dig out roots and tubers. Also during courtship
duels the animal can put to the opponent stings. Jaws of brontothere hornhog
are long and rather weak. The basic food of animal includes young grass and
leaves of bushes.
The brontothere hornhog is able to run quickly and long – at it legs are rather
long. In this connection hoofs of this animal are small – this way friction
against the ground is less. But it imposes the certain difficulties: the animal
is compelled to avoid fenny areas, and comes into water only in case of emergency.
Animals avoid places where there are flooding. Differing in ability to predict
weather, they leave lowland areas to which flooding may threaten, and migrate
to foothills. At this time separate family groups may unite, and then between
males fights happen.
This species keep in small herds (up to ten adult animals), each of which represents
a harem of one male. Animals communicate with each other by means of various
sounds uttering with the help of short proboscis (at this moment it is slightly
puffed). The young growth keeps with parents before independence then males
are expelled by the leader, and females – by adult females. Seasonal prevalence
in breeding is expressed poorly, but the majority of cubs is born in the beginning
of rain season. In litter there is always one cub.
Bullhog
(Buchoerus crassoceras)
Order: Even-toed ungulates (Artiodactyla)
Family: Horned hogs (Porceratidae)

Habitat: savannas and light forests of Southern Africa, damp sites near to
rivers and bogs.
When during evolution process in nature a “successful” variation of live beings
appears, on its basis related forms adapted to different conditions of inhabiting
at once start to evolve. In Southern Africa at the basis of the common ancestor
different species belonging to new Neocenic family of horned hogs were evolved.
These animals have partly replaced large ruminant artiodactyl animals, not
sustained chasing from the side of people and destruction of habitats during
a catastrophic congelation. Hornhogs became analogues of buffaloes, large antelopes,
and partly even rhinoceroses. Bullhog is the largest representative of family.
This species of mammals differs from brontothere
hornhog by more powerful addition: the animal is very similar to large
buffalo. The highest point of the body is at shoulders, back is a little bit
sloping. Wool is short, pale-yellow, and lighter on stomach. On cheekbones
wool forms black “whiskers” especially obviously expressed at males. Legs,
partly shoulders and groats of animal are covered with short horizontal strips
of rusty wool. Tail is long, with hairy brush on the tip.
The most characteristic attribute of animals of horned hog family is the horns
of various shapes developing in middle part of head. At bullhog, despite of
the name, horn it is completely not similar to bull horns: it is wide and flat,
reminiscenting somewhat the horn of living in North-African savannas flathorn
(Platyceratherium foetidus), huge animal from order of damans (Procaviiformes).
It is wide, covered with cornificate skin, and concave in middle part. The
horn of bullhog is mainly the tournament weapon, rather than fighting one:
in courtship season between males of this species there is a force struggle
by necks (muscles of neck are very strong). Horns serve as protective “helmet”:
they are formed by thick layer of bone tissue with a minimum quantity of cavities.
But also this horn may be used as the weapon: to attacking predator protecting
bullhog puts ramming impact by head. The horn protects eyes from casual wounds,
therefore, having “hooked” predator by flat horn, bullhog can lift it in air,
and then simply strike a predator against tree or throw off far.
As against the relative, the brontothere hornhog, bullhog prefers sites which
may be flooded. This animal willingly bathes and wallows in dirt, and also
searches near to reservoirs for significant part of food – soft vegetation.
In this connection at bullhog the wide hoofs able to move apart partly, when
the animal grazes at the bog, were developed. Also the animal is able to swim
very well, and easily crosses even wide rivers and lakes.
Bullhogs keep in small harems: one male and 3 - 4 females with cubs of the
age till one year. Such harem occupies the certain territory along the river
bank, or in bogs, and does not leave it even at flooding. Except for such families,
there are single males living at the narrow border lands between possessions
of family males. Usually they are either very old males expelled by young contenders
from own groups, or young males, yet not having own harem. In courtship season
young animals challenge family males, and between them there are skirmishes.
Once a year the female gives rise to 1 – 2 cubs. They differ from adults by
darker colouring in which there are more strips (they stretch even on back
and sides). The female carefully preserves them against predators, and easily
may attack any animal if it will seem that the invader represents threat for
the cub. The young growth is fed with milk till four months, and then completely
passes to vegetative forage. One-year-old youngsters abandon herd: shortly
before birth of the new cub the female starts to show aggression to them.
Sexual maturity comes at the age of about 3 years, life expectancy may account
till 40 years.
Hyena-like
dienaper (Deinaper crocutoides)
Order: Even-toed ungulates (Artiodactyla)
Family: Predatory boars (Carnoporcidae)

Habitat: grass plains and bush thickets of Central and East Asia.
In Paleogene and early Neogene among characteristic inhabitants of open spaces
of Eurasia and North America there were entelodons (Entelodontidae) – family
of non-ruminant artiodactyls, related to pigs. By structure of teeth, they
were omnivores in which diet meat took an essential place. After epoch of mass
extinction when many ecological niches appeared free, pigs have taken advantage
of a situation, and have taken a vacant place of scavengers. So at plains and
in bush of Asia the family of Predatory boars has partly occupied a place of
hyaenas and bears has appeared.
The typical representative of this family is deinaper (“deino-“ means “terrible”,
and “aper” – the boar), the largest species in family. An ancestor of this
animal was wild boar (Sus scrofa), the species widely widespread in Eurasia
and have successfully survived the period of anthropogenous pressure. The important
place in diet of boar was occupied with food of animal origin. One of the evolutionary
lines, originated from this species, has gone to the way of adaptation to searching
and eating larger amount of food of animal origin. Gradually these animals
have turned to scavengers, and even had become able to attack cubs of other
animals.
Deinaper is an animal of massive constitution, its growth at a shoulder is
about 0.8 m and weight is up to 140 kg. By constitution it is similar to the
spotted hyena: at it there is large head on mobile neck. Its body is covered
with shaggy rough wool; to winter rich underfur grows. In connection with unusual
for hoofed mammals way of life at it spotty colouring developed – strips characteristic
for boar juvenile colouring, break up to separate spots. It even more strengthens
similarity of deinaper with hyenas. Legs of animal are rather long, adapted
to fast running to the big distances. Because of long legs the body seems a
little bit short.
Jaw are short and high, able to crush bones. There are large sharp tusks jutting
out up and in sides. They are used as the weapon in tournament fights, but
in case of need deinaper can use them against predators.
The appearance of muzzle of deinaper is characteristic for pigs. Sense of smell
is sharp - with its help animal searches for carrion and finds small animals.
Deinaper is aggressive: it can drive away medium-sized predators from catch,
threatening them with tusks and making sharp attacks on aside a predator. As
this animal lives small groups (up to 4 – 6 adult animals), not any predator
can defend the right to the catch in the face of several strong and aggressively
adjusted animals.
The group of deinapers supervises extensive territory on which there are some
shelters. Borders of territory are marked by heaps of manure, and constantly
renewed. In shelters animals spend hottest time of day, becoming more active
in the morning and in the evening when the majority of predators hunts. If
won over catch at predators does not suffice, deinapers can hunt in the afternoon.
Then they attack small animals having a rest in a shadow.
Once per year the female brings posterity: up to 5 – 6 cubs. Feeding female
does not take part in hunting, protecting posterity. The only thing, that it
presumes to herself – to catch any tiny casual catch like reptiles and rodents.
But males and females without cubs after successful hunting necessarily belch
for feeding female pieces of meat. Newborn cubs are weak and helpless: they
start to stand only at the age of one week, and pass to feeding by meat only
at bi-monthly age. Approximately at the age of three months young animals leave
lair, and lead nomadic life with parental group. Only at the second year of
life the young female can give posterity the first time. And the young male
can really apply for domination in clan only since the fourth year of life.
Up to this time it leads life of the lonely hunter in boundary territories,
gaining strength and experience to challenge once in winter to any old leader
of clan and to take his place.
(This animal is discovered in common: by Tenek, the participant of forum (she has stated the assumption of existence of the present animal in Neocenic fauna), and by Arseny Zolotnikov (he created initial sketch of the present animal))
Armor-headed
wart hog (Geophacochoerus subterraneus)
Order: Even-toed ungulates (Artiodactyla)
Family: Pigs (Suinae)
Habitat: savannas and light forests of Southern Africa.
Ancestor of present species is usual in Holocene African wart hog (Phacochoerus).
These hoofed mammals spend significant part of day in holes dug independently.
And its descendant has finished this habit of life to extreme being shown on
ground surface only at night. This animal spends day in shelters – wide holes
at times directed downwards to two meters depth and stretch under ground to
tens meters. In connection with such original habit of life the shape of animal
had strongly changed.
The armor-headed wart hog reaches height only 40 cm at a shoulder. But its
body is very much extended – almost meter longwise not including tail. About
20 % of body length the head turned to tunneling mashine amounts. At this animal
there are large, slightly bent tusks sticking up forward and in sides – this
animal digs and deepens holes, at feeding on surface overturns stones in searches
of insect larvae and digs out plant tubers and rhizomes by them. At the top
part of head thick layer of cornificate skin protecting head from casual fallings
in holes is advanced. Tip of muzzle with “snout” characteristic for pigs is
very mobile that partly compensates hindrances created by tusks at feeding.
Nostrils of animal are closing during digging. Nasal courses are rather short
but the olfactory surface forms numerous plicas: at armor-headed wart hog it
is very sharp sense of smell. Sight of animal is rather weak; eyes are protected
from ground by dense eyelashes. Ears are small; at digging they fold lengthways
and nestle to sides of head. On cheekbones the large bone outgrowths especially
advanced at males stick up.
Body is long and legs are short. On forward legs in front, and on back ones
under hock there are calloused outgrowths serving as a support at ground digging
(animal digs a hole standing “on knees”). Hooves are wide; at digging they
are used for shoveling up the dug ground. Hairs on body are submitted by short
thin bristle, “mane” on back characteristic for wart hog is not present. Skin
is wrinkled and grayish-black. At the tip of long tail there is brush of white
hair: tail is used for communication and signal submission.
Armor-headed wart hogs live in groups: dominant male, female and cubs of current
litter (up to 5 cubs 2 times per one year are born). At new litter birthing
at the female grown up young animals are driven out. The family of animals
lives in constant territory which is jealously preserved against neighbours.
At the territory usually there is a main hole and plenty of temporary refuges
well known to all family of hogs. In due course holes are extending, sometimes
systems of holes join to each other. At exhaustion of food sources around of
main hole the family moves (at night or dawn) to other hole, somewhere to the
edge of territory. Gradually the centre of territory is actually displaced
here that inevitably results to revision of borders between neighbours.
In each hole there is a main chamber where the female with posterity lives,
and “throne hall” where the dominant male lives. They are located opposite
to each other and between them main burrow with two exits (one of which is
disguised outside by dust) passes through. On each side of this pass animals
dig out temporary toilet chambers leaving dung there. After filling the chamber
is filling up by ground and simultaneously the new chamber is making. This
way animals hide their presence from predators.
In holes of armor-headed wart hogs a plenty of “lodgers” settles: butterflies,
crickets, numerous fleas (they parasitize in thin wool on throat and stomach
of animal). Among vertebrates large tortoises and lizards (hogs simply can
to have eaten small reptiles) and also birds and small mammals settle in holes.
For protection against parasites wart hogs willingly take dust baths sometimes
arranging place for bathing right in heap of ground thrown out from hole.
Family of armor-headed wart hogs goes for feeding at night. At first male appears
from hole; he looks, listens and smells around determining whether there is
no danger. If all is quiet he walks out and after him female and cubs appear.
At night animals go to watering place (if the water source is far some burrows
to it are dug allowing to not show itself to superfluous danger at the watering
place) and then feed on meadow or under trees. The basic forage of armor-headed
wart hogs is grass. But in trees and bushes fructification season they willingly
eat drop fruits and berries. Animals willingly eat forage of animal origin:
worms and beetle larvae gathering them at hole digging, searching under stones
or in rotten wood. Differing in keen sense of smell such pig easily feels presence
of larva under thin layer of wood and if it is necessary splits mouldering
tree by sharp tusk. If it is possible to find carrion or remains of predator’s
catch animals willingly feed with meat.
During feeding armor-headed wart hogs communicate wagging with white hairy
brush on the tail tip. Their voices at this time are similar to usual pork
grunt. Tail sharply lifted upwards is an alarm signal: having seen such mark
animals cease eating and start to smell around trying to find out predator.
In case of danger animals squealing shrilly rush to nearest hole trying to
keep near it at all. These pigs are fast and maneuverable runners though they
can not run long on short legs. At deviation male climbs to the hole at last
having turned to predator by head. He loudly roars trying to frighten predator
and from time to time makes aggressive lunges trying to strike attacking predator
by tusk. Usually after that only a little number of wishing to continue chasing
ones may be founded.
Maturity at females comes at two-year-old age, at males little bit later. Young
females at the second year of life pair the first time and approximately after
three and half months give rise to posterity. The first litter is insignificant:
no more than three pigs and usually only two ones. But at the fourth year of
life in litter it may be up to five pigs.
Young growth abandons parents shortly before birth of next cubs or right after
it. Youngsters first time keep in parental territory using for life old holes
- at them tusks are still insufficiently grown. But when the young growth new
litter grows up, the dominant male, their daddy, expels all of them from territory.
At this time young animals perish in a plenty from predators. But ones managed
to find suitable place for life, have an every prospect to live up to old age:
the age limit of these animals reaches 15 years.
Deersimil
(Cervops platycornis)
Order: Even-toed ungulates (Artiodactyla)
Family: False deer (Neocervidae)
Habitat: savannas and light forests of Southern Africa.
After extinction of significant part of large hoofed mammals their place was
occupied with representatives of other groups of mammals: in Palearctic there
are large running descendants of lagomorphs, and in Nearctic and Neotropic
areas – descendants of South-American rodents. Only south of Africa appeared
rather isolated from the Sahara savanna by woods and mountains. Here again
species of herbivorus mammals have remained relicts of Holocene dominant: various
hoofed mammals, descendants of small antelopes and chevrotains.
Occupying ecological niches exempted after mass extinction they turned to species
similar to well-known in human epoch antelopes and deer. At some species even
big horns have appeared – it is a rarity among Neocaenic herbivores.
The deersimil is not the most numerous species among South-African hoofed mammals.
It keeps in small herds sometimes together with other species of herbivores.
This animal is the descendant of small and short-legged African chevrotains
(Tragulus). Having remained out of competition from the side of bovines chevrotains
roughly have started to evolve having formed the separate family of false deer
– rather large swift-footed hoofed mammals.
In comparison with ancestors deersimil is a sizable animal: its growth at a
shoulder is up to 1.5 m, body length is up to 1.6 m. By constitution it resembles
small antelopes and gazelles. This is one of most swift-footed hoofed mammals
of Neocene: at short distances deersimil accelerates momentum up to 80 kms
per hour. Because of long legs usual pace of this animal is an amble but deersimil
develops maximal speed running gallop.
Colouring of deersimil helps it to hide among vegetation: body is covered with
chestnut-brown wool; on groats and hips there are thin vertical strips of yellow
color. Stomach, bottom part of neck and head sides are white. Cubs are colored
lighter: the basic colouring at them is straw-colour with longitudinal brown
strips on body (by colouring they are similar to boar piglets); on groats and
hips strips are bent downwards. In process of growth striped pattern on the
body vanishes and on hips becomes more contrast.
This animal has horns on head. But as against deer horns deersimil horns are
constantly growing (not dropping as at deer), shovel-like and with several
jags (“antlers”). These horns are a little similar to horns of African prehistoric
giraffes Sivatherium. Bases of horns are enough strong and extended along of
skull; horns grow on head of animal in parallel to each other. Horns of females
resemble more longitudinal crests with two jags at back edge. At males horns
are larger and it can be up to 5 - 7 jags on them; horn tops at males slightly
inclinate in sides. New jags appear on forward part of horn: at old males horns
are sometimes similar to rooster crest. Similarly to deer deersimil uses horns
as the tournament weapon - opponents approach to each other so that horn jags
get in dredgings on contender’s horns. At the duel basically a power struggle
takes place – contenders try to push aside and tumble down each other.
From senses at deersimil hearing and sense of smell are most sharply advanced.
Other herbivores use it watching for behaviour of these animals for detection
of probable danger. In case of alarm the deersimil utters special shout - sharp
whistle warning of neighbours and other animals grazed near about danger. Rescuing
from predator deersimil hopes for the speed in many respects - few
predators can catch up adult animal full of forces. Even two-day cubs are
already almost equal in speed to adults though, certainly, they get tired much
faster.
Deersimil eats mainly graminoids. Grazing animal bites off only their tops;
after feeding of this animal other species of herbivores can find enough food
at the same place to themselves. Especially deersimil likes cones of graminoids
in condition of wax ripeness: thus some graminoids differ in sweetish taste.
Usually deersimil feeds during dawn and sunset and spends hottest time of day
in tree shadow.
For protection from overheat deersimil uses the original adaptation – its own
horns. As against deer horns they are always combined by alive growing bone.
Outside of breeding season skin on horns is plentifully penetrated with blood
vessels serving as fine radiator: blood emits heat to air. But in breeding
season because of hormonal influence blood vessels in basis of horns are narrowed
and substantially block stream of blood to horns that protects males from blood
loss at casual wounds.
Breeding season at deersimils begins at the end of rain season. Between males
there are short skirmishes for “harem” of several females (depending on physical
condition male can declare rights to 4 – 10 females). Opponents usually avoid
to enter direct fight and all can be limited to demonstration of force: males
bellow bassy, dig ground by hoof, widely open mouthes and show teeth. Sometimes
they rear and showing to each other the might make some steps on rear legs.
Usually one of contenders leaves stadium without fight.
Time of pairing proceeds only about one week and then aggression of males is
reduced also they become tolerant concerning to each other again feeding sometimes
side by side.
After dry season female gives rise to one or two cubs (about half of females
fawns twins). As at all hoofed mammals cub is born advanced and in some hours
it is capable to follow mother. First two months it feeds by milk but from
fortnight age it already tries plants copying mother. To the end of rain season
it becomes independent, but stays in herd. Sexual maturity at females comes
at the age of one and half years, at males at two years.
Sunhorn deer (Heliocervus heliocornis)
Order: Artiodactyls (Artiodactyla)
Family: Deer (Cervidae)
Habitat: eastern part of North America, light forest and bushes in plain and
foothill areass of Appalachian Mountains.
Till whole human epoch, since prehistoric times, deer were traditional object
of hunting. Chasing of these animals had resulted in their preservation only
in few places of their former area. In Northern hemisphere these places were
remote northern grounds, Siberia in Eurasia and area of Rocky Mountains in
North America. After the ending of human epoch some small deer species preserved
in forests of Southeast Asia. These animals became ancestors of new species
of deer of Neocene epoch.
North American deer are descendants of white-tailed deer (Odocoileus virginianus)
– one of the most usual species of the family at this continent. One of descendants
of this species is magnificent sunhorn deer. It is medium-sized deer – adult
male weighs about 100 kg, female is much smaller. For this one gracile constitution
is typical – this deer is able to run quickly. Sunhorn deer lives in light
forest areas where bushes alternate with sites overgrown with grass. The basis
of ration of this animal is made of leaves of bushes and low trees and also
of rather soft grasses not belonging to graminoids.
The body of sunhorn deer is covered with wool of bright yellow color; dark
longitudinal strip stretches from the nape up to the basis of tail. Occasionally
at separate individuals in adult condition attributes of juvenile colouring
keep – white spots on shoulders and hips. Stomach at females is white. At males
colouring of stomach is quite another – it is white with numerous black spots.
This colouring is used by males for display in courtship season. The bottom
part of tail at individuals of both genders is white. Vertically rised tail
making seen a wrong side appreciable from apart is the traditional alarm signal
at deer.
Deer are an excellent example of action of sexual selection. Various species
of these animals during the history of this group developed horns of various
sizes and shapes. Extreme degrees of expression of consequences of action of
this kind of selection are fossil deer Megaloceras with huge horns and Eucladoceras
on which fan-shaped horns more than ten antlers developed. Sunhorn deer had
actually repeated an evolutionary way of fossil deer Eucladoceras – its horns
have completely lost function of the tournament weapon, and have turned to
object of courtship display. The shape of horns of this deer has determined
its name. Horns of sunhorn deer have set of antlers which grow practically
in one plane. In the basis of such horn there is wide semicircular “shovel”
as at the elk, but turned vertically. On its edge numerous straight antlers
grow, because of what both horns of this deer create the impression of stylized
solar disk with beams. At adult males it may be up to 8 – 11 and even more
antlers on each horn. In courtship season males avoid combat, being limited
to displaying of horns. Females of this species are polled; it is typical condition
for all deer, except for odd-looking skewhorn from Greenland.
Within summer season males of sunhorn deer concern to each other tolerantly
and even form barchelor herds numbering up to 20 individuals. The courtship
season begins in the middle of an autumn. At this time horns completely ossificate,
the rests of skin fall down from them, and deer males become aggressive. Males
of this species gather at “stadia”. They display themselves to contenders,
rearing. Keeping balance, male can make too many steps in vertical position.
Thus spotty colouring of its belly becomes especially good seen. Usually males
compete; they try to keep longer a vertical position – this way the strongest
one is revealing among them. If rivalry is critical, males can pass to active
struggle against contenders: they push each other by chest, trying to tumble
the contender down. Occasionally they strike impacts to each other with the
help of forelegs.
The winner male gathers a harem of 5 – 8 females and within several days repeatedly
couples with them. He furiously rushes to every one, in which it sees the contender
(even to animals of other species), and during the courtship season eats almost
of nothing. To the end of courtship season male is strongly exhausted, and
some males even perish, including from a nervous exhaustion. In winter males
mew horns.
Pregnancy lasts about half-year, and cubs (usually twins) are born in spring.
Juvenile colouring of sunhorn deer is spotty: cub is darker, than adult individual,
and on brownish background of its wool white spots are scattered forming faltering
longitudinal strips on back and sides.
To an autumn young deer leaves mother, and at this time she is ready to new
pairing. At the second year of life young females take part in breeding season
the first time. Males can take part in courtship tournament only since the
fifth year of life. For the second year they grow primal small horns which
have few antlers and no “shovel”. Each new year of life horns become larger,
and at five years get the shape characteristic for horns of adult animal.
Maned spirocervus (Spirocervus jubatus)
Order: Artiodactyls (Artiodactyla)
Family: Deer (Cervidae)
Habitat: Himalayas, mountain forests.
In human epoch the majority of large animals had been exterminated as a result
of hunting, or their areas and number had been strongly reduced because of
destruction of habitats. As a result after human disappearance the majority
of large animals had died out, not having left descendants. Their place was
occupied by large descendants of more successfully survived small species of
animals.
Asia was the most densety populated continent of human epoch. This circumstance
became the reason of practically general degradation of natural ecosystems
and extinction of large animals. But in Neocene this continent appeared populated
by large animals again. One of such species is the descendant of small barking
deer (Muntiacus muntjak) which lives in Himalayas. This animal is equal to
large deer of human epoch in size (height at a shoulder up to 120 sm, weight
over 150 kg) and is the approximate analogue of bezoar goat of human epoch.
Similarity is strengthened even more by the shape of horns of this deer – they
have no characteristic branchy shape and are mostly straight with spirally
twirled edges. This feature has determined the name of animal – spirocervus
(literally “spiral deer”).
Spirocervus has robust constitution; its rear legs are little bit longer than
front ones, and back is inclined forward. It is the prominent feature of herbivores
living on hillsides – it is easier so to graze. Also at spirocervus there are
strong hooves behind which the elastic small pad is located. Hooves can easily
move apart. Due to such features of structure of legs the animal is able to
move on abrupt stony slopes though prefers to live in mountain forests.
The high mountains differ in rather cold climate. The alternating of seasons
is clearly expressed here, and it is cold weather till the significant part
of year. Spirocervus is adapted to such features of mountain climate. The thickset
constitution reduces a relative surface of its body. Brown colored wool of
this animal is long and rich. In cold months it becomes thick, but does not
change color, as at snowlopper from the number of hoofed lagomorphs, living
higher. On breast and neck of spirocervus males long mane grows, and tail is
ended with magnificent switch of white color.
Head of spirocervus has odd appearance. On head of this animal two longitudinal
bone crests from nostrils up to the bases of horns grow – it is a characteristic
attribute of barking deer descendants, including massive bulldeer of China
swamplands. At the female bone crests are less expressed, and horns are lack.
And horns of the male have very unusual shape. They consist of two antlers
– the basic one growing upwards, and lateral one directed aside and outside.
The basic antler is straight; at the adult male it reaches the length 70 –
80 cm. It is three-edged in cross-section and is slightly twirled spirally.
Lateral antler is short, thick and hook-like. It serves for fixing horns at
courtship tournaments.
Bone crests pass to bases of horns. They are covered with long wool of yellowish-brown
color at males and grey at females. At males wool on bases of horns is so long,
that it seems the continuation of mane.
Spirocervus precisely differentiates habitats with snowlopper. It avoids places
where snow lays till the most part of year, and does not meet higher in mountains
where the snowlopper lives. Usually spirocervuses keep in herds of 10 – 15
animals – females and young animals of both genders under leadership of the
adult male. Sometimes in large herd it may be two adult males. These deer live
in mountain forests and feed on leaves of bushes and low trees. In winter when
leaves fall from most part of plants and forage reserve of animals is reduced,
spirocervuses may eat leaves of evergreen rhododendrons which they usually
do not touch in summer.
In the beginning of an autumn the courtship season begins. At this time males
roar loudly, declaring rights to territory and females. Bachelor males (usually
these ones are young animals) challenge males, interfering to their territory
and roaring. Between competing males there are severe duels for harem. They
are reduced to force struggle of animals linked by horns. Lateral antlers on
horns serve as terminators, not allowing animals to injure each other; therefore
males usually go through this time without damage.
Pregnancy at the female lasts about half-year, and she gives rise only to one
well advanced cub. Such tactics is more favourable in conditions of limited
resources. Cubs of spirocervus differ from adults in dark, almost black colouring,
with several small white spots on throat. In the course of time they brighten,
and the wool changes shade to brown color. To winter the young animal becomes
almost independent. Females are ready to breeding at the third year of life.
At males at the second year of life simple forked horns with almost equal basic
and lateral antlers grow. Next years horns change the shape, in brief repeating
an evolutionary way of ancestors of this animal – vertical antler becomes longer,
and lateral one grows thick.
Life expectancy makes about 40 years.
Bald rambull (Pachycephalovis grandis)
Order: Artiodactyls (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: Himalayas, Plateau of Tibet, Altai; mountain woods and meadows at
height up to 4 – 5 thousand meters; in the north of area animal goes down,
but does not meet in plain regions.
At the boundary of Holocene and Neocene the variety of ungulates had sharply
reduced. The reason of it had been mainly human activity/ People had overpopulated
the planet and had destroyed habitats of many species. Agrarian landscapes
changed by people, where herds of domestic cattle were analogues of wild animals,
became the prevailing landscapes in many places of the Earth. After human extinction
the majority of domestic animals had also disappeared, but some species able
to adaptation, had survived and became ancestors of new forms of Neocene ungulates.
Among them there were primitive breeds of domestic sheeps which easily ran
wild and became a part of poor communities formed at the place of collapsing
agrocenoses. Many descendants of domestic sheeps live in Neocene New Zealand,
but separate species had evolved in other parts of the world.
In mountain areas of Asia large artiodactyl – bald rambull is found. Height
of this animal at a shoulders is up to 170 cm at body length of about 2,5 m.
Externally it is similar to golden goat antelope (Budorcas taxicolor), the
original representative of bovids, related to sheeps, lived at the territory
of China. The body of rambull is covered with rich shaggy wool of gray-blue
color, on back the longitudinal strip of darker wool stretches; the tail is
covered with long wool and is similar to horse’s one. Ears are moderately long
and covered from external side with wool.
As against New Zealand sheep descendants, bald rambull has almost completely
lost horns. Instead of it the upper part of skull at this animal is strongly
thickened. The skull of male has the convex top representing a continuous bone
outgrowth about 15 cm thick. Males use skull for courtship tournaments during
which they put each other strong ramming impacts. Thus the massive cover of
skull protects brain of animal from concussion. On top of the head of animal
the skin is dense, thick, strongly cornificated, colored dark grey, completely
lack of hairs. Near ears of animal there are outgrowths hided in wool, also
covered with cornificated tissue – they are rudiments of horns. Females lack
of them more often. Also at females the top of skull is not as convex, as at
males, and they do not use it for fierce tournaments. Only occasionally females
strike each other easy impacts by head, reminding the subordinated individual
about their own position in herd hierarchy. Cervical vertebrae are adapted
to maintain the sizeable loadings.
The thick ossificated top occupies the most part of volume of skull. The brain
of bald rambull is very small, and the behaviour of animal differs in primitiveness.
Weakness of sight is compensated, however, by keen hearing and sense of smell,
and also by natural care. Animals move across their territory along strictly
determined routes, and only natural acts like avalanches or earthquakes force
them to search for new ways. This species of ungulates lives in small herds
including one dominant male, several females and their cubs of the age of up
to two – three years. In herd the not strict hierarchy is established.
Bald rambull eats leaves of undersized trees and bushes. At lack of habitual
forage it can eat even branches of coniferous trees. Also the animal is able
to dig snow in searches of grass and evergreen bushes. This species lives in
conditions of seasonal climate – winter in habitats of bald rambull is frosty
and snowy. Therefore right before colds it grows long rich wool of lighter
color, rather than summer one.
Pregnancy lasts 11 months; usually the female gives rise to one cub. In favorable
years twins are often born – one case to approximately 8 – 10 pregnant females.
Cubs are fed on very fat milk and quickly grow. At the age of about 8 months
the young animal completely passes to forage of adult animals. At young animals
approximately up to one-year-old age the head is covered with wool, as at other
representatives of family. At the age of four years animals completely develop
physically, and females already can bring posterity. Young male reaches the
blossoming of physical strengths to the seventh year of life, and supports
this physical condition at least till 15 years.
New
Zealand orovis (Orovis austro-alpinus)
Order: Even-toed ungulates (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: mountains of New Zealand from wood zone up to Alpine meadows.
New Zealand during millions years had remained “the lost world” even more forgotten
rather than South America of Paleogene and Neogene epochs. There were no even
mammal species there except for pinnipeds for which ocean open spaces are not
a barrier and bats have arrived to these islands by air. People have thoughtlessly
“enriched” unique fauna of islands with various species of domestic and exotic
mammal and birds. Thus catastrophic damage was put to local fauna.
After human disappearance on Earth New Zealand has remained in any measure
to the “lost world” condition separated from other world by Pacific Ocean.
Animals belonging to groups dyed out or fairly degrading in “big world” had
kept here. So, when number of hoofed mammals on continents had reduced, in
New Zealand one of their species had roughly evolved. Pacific Ocean by its
heat had kept this ground from congelation and has provided enough precipitations
for grass and trees developing on these islands. And in Neocene descendants
of one of survived species, domestic sheep, wander among meadows and woods.
There are swift-footed wood animals, massive inhabitants of plains and light
forests, and also dexterous and mobile inhabitants of mountains among them.
New Zealand orovis (literally: “mountain sheep”) has returned to habitat where
distant ancestors of these animals lived: in mountains. Here, on Alpine meadows,
small herds of these animals under the leading of dominant male graze.
Orovis is medium-sized animal (domestic goat-sized one). Back legs at it are
longer than front ones: it is a characteristic feature of mountain animals.
At such constitution it is more convenient to graze on slope.
Neck of animal is strong especially at the adult male - animals establish hierarchy
butting. Horns are short, straight, with thick bases, sticking up back and
in sides. Forehead is very wide, at the male it sometimes acts as osseous “helmet”.
Muzzle of orovis is narrow and rather long: it helps to nibble grass growing
between stones.
Hoofs of orovis are strong and wide. The bottom surface of hoofs is concave
and edges are rather sharp: with the help of such hoofs animal can walk on
abrupt slopes and keep on smooth rocks.
In mountains it happens coldly enough even in warm climate of Neocene. Therefore
orovises differ in dense and thick wool. Adult animals have grey wool on the
body, dark legs and “belt” (longitudal stripe) on back. For recognition of
neighbours on cheeks of adult animals there are white stains varying by size
and form at different individuals. In summer animals fade and wool becomes
shorter and thiner.
Summer is time when air rings from flights of different blood-sucking flies.
Therefore tail of orovis had turned to excellent fly-beater – it is long and
flexible with brush of long hair on the tip.
Orovises migrate in herds in mountains of New Zealand. In each herd there are
large dominant male (large head with thick horns distinguishes it), females
standing lower in hierarchy (in harem of one male there are about ten ones
of them) and cubs. In summer these animals rise highly in mountains, and at
times they reach snow-line. Their basic food is grassy plants and mainly graminoids.
It is much more difficult to search for forage in winter – orovises hardly
rake snow with their thin legs. Therefore more often in winter it is possible
to meet orovises in foothills where snow layer is not so thick or it does not
fall absolutely. Animals feed in woods, graze grass and ferns, and gnaw branches
of bushes and young trees. At this time some harems of different males can
unite: in winter these animals tolerantly concern to each other. Also it is
easier to big herd to protect from local predators.
In spring when snow in mountains thaws orovises come back in mountains. At
this time between males there are fierce duels for females and territory: opponents
ram foreheads and make force struggle trying to tumble the contender down.
Both contenders loudly roar during a duel trying to make an impression upon
the opponent. Sometimes in the heat of struggle they rear and start to beat
opponent by forelimbs. After such duel defeated one runs out from stadium but
sometimes at worst it stays on a battlefield lying having had neck fracture.
The winner male drives “harem” of defeated one to the herd and marks new territory
by dung heaps. The defeated male now stays practically without means of subsistence:
it lives at the edge of territory which once was its property and carefully
hides the stay on it – up to the following breeding season.
Once a year in early spring while animals stay in forest zone, the orovis female
gives rise to two cubs. They can run in some hours after birth. When snow in
mountains thaws the herd goes to Alpine meadows and during the migration cubs
master elements of mountaneering. They suck milk up to 4-month age but since
the second week of life start to try plants eating by adult animals. Grown
to the age of half-year young males start to suit duels and to establish hierarchy.
At this time dominant male starts to show them the superiority compelling them
to leave from herd.
Young males form independent herds and migrate in mountains within two years.
Having reached maturity they start to challenge adult owners of “harems”, and
at times the herd of old male appears shared between two or three young applicants.
Taurovis
(Taurovis aotearoae)
Order: Even-toed ungulates (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: marshy plains and bushes of New Zealand in temperate climate zone.
Up to time of European colonization of New Zealand first settlers of these
islands had already time to destroy huge wingless moa birds occupying ecological
niches of herbivorous mammals. Ecological niches which have become empty as
a result of unlimited hunting of aborigenes stood empty rather not for long
time: after disappearance of humans their satellites – become wild pets – have
occupied them. Descendants of domestic sheeps, fertile and quickly adapting
animals have occupied niches of herbivores in various places. One of sheep
descendants had turned to very remarkable animal resembling bulls by mighty
constitution.
Taurovis, the “bull-sheep” – it is the name of this creature. This is very
large animal, largest one among New Zealand herbivores: its height at a
shoulder is up to 1.7 m at length of body up to 3,5 m. At it there are massive
body, rather short thick legs and large head. The neck of animal is short and
covered with mane of long hair. The body is covered with short brown wool with
dark “glasses” around of eyes and “stockings” on legs. At males wool on trunk
is darker than at females.
The “ram surface” of head of the mature male is trapezoid, flat, formed by
forehead and wide bases of short thick horns and covered with hairless cornificate
skin. Ends of horns are directed to sides and even a little downwards. Horns
serve to these animals rather for demonstration than for direct purpose: it
is impossible to make force struggle by them, at butting horns of animals do
not adjoin at all. At taurovis females horns are underdeveloped, and look as
bone lumps on each side of skull and forehead is more convex than at males.
Edges of forehead hang above eyes forming original osseous “peaks” protecting
eyes from casual damages by branches of bushes.
Taurovises live on bush plains of New Zealand frequently coming to bogs. For
walking across fenny bogs this animal has special adaptation: hoofs are wide
and can move apart increasing the area of support. Besides these animals perfectly
swim and frequently feed in rivers thining out thickets of reed and other marsh
plants. In heat animals spend all day in the river grazing on bank in the evening.
Near rivers one trouble waits of these large animals: in water mosquitoes,
midges and other blood-sucking insects breed in plenty. They attack warm-blooded
animals by large swarms exhausting them by stings. Taurovises are able to resist
to this mistfortune: they bathe in river or wallow in dirt. The drying up dirt
unpleasantly smells partly masking smell of animals. Besides it sticks wool
of animals to the true armour impenetrable for stings of mosquitoes.
In places where taurovises regularly feed bushes do not form continuous cover:
big animals trumble and constantly renew tracks along which they walk. It expands
their forage base: among bushes clearings appear where graminoids and other
various grasses giving food to taurovises and other local herbivores plentifully
grow.
Taurovises live in herds including one male, some females and cubs of first
two years of life. Each herd has the territory which borders are marked by
manure heaps. These heaps are regularly renewed, and at times reach 1 meter
height. Herd has some basic routes of movement on territory. Within one year
they are used with different frequency: in summer animals spend time near to
rivers or lakes, and in winter more often in bush thickets.
Once a year female gives rise to one large cub. At three – four hours after
birth it stands and tries to walk, and for the second day of life it freely
walks behind herd. Young growth is covered with more light wool than adult
animals: it is straw-coloured with brown legs. Young taurovis spends first
two years of life in parental herd. But when it grows up and colouring of wool
becomes all more similar to adult one, young animal is compelled to abandon
herd – adult animals show aggression to young ones seeing competitors in them.
Young females give rise to posterity since four-year-old age.
Sylvammon
(Sylvammon gracilis)
Order: Even-toed ungulates (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: evergreen woods of Northern island of New Zealand.
Forests of New Zealand had lost because of human destructive activity large
moa birds in Neocene had found new herbivorous inhabitants. Descendants of
sheeps, domestic animals introduced to New Zealand became them. In Neocene
when the human species had disappeared and set of new ecological niches had
appeared descendants of this domestic animal had started to evolve roughly
having evolved in conditions of island isolation to some species differing
by ecology.
The sylvammon is large species of artiodactyls living in woodlands and forests
of New Zealand. Withers height of adult male is up to 150 – 160 cm. By proportions
this species is a little similar to deer.
The long neck is covered with mane of lengthened hairs. At old males mane hangs
down almost up to knees of forward legs. Shoulders of sylvammon are higher
than crupper and back of animal is sloping. Due to such constitution animal
can browse leaves from trees at height up to 2 meters. Due to need animal can
even rear on back legs. Animal can run quickly and jump high.
Horns are similar to goat’s ones; at females and young males they are straight
and only at old males can bend semilunarly. Bases of horns are bulged and also
pulled together, and horn tips are directed back – in this case they do not
prevent to move in dense wood. Among females hornless animals appear frequently.
Horns at males are lighter than at females.
Wool of sylvammon is short and rather thin: animal lives in rather warm climate
not rising highly in mountains and not coming to southern areas of New Zealand.
Except for mane long hairs grow on tail of this animal more similar to horse’s
one. Colouring of body of sylvammon is dark-brown with white stomach and white
“glasses” around of eyes.
It lives in woods preferring to graze at sites where bushes and young trees
spread off: on marges and near river banks. Frequently it goes to plains overgrown
with bush. In wood animals have favourite tracks adhering them at movement.
Sylvammons have kept characteristic for sheeps gregarious habit of life. Herds
of these animals total up to 20 individuals and consist of several breeding
groups including male and three – four females. In forest herd moves having
stretched in line where ahead there are dominant individuals. The rank of females
in herd is determined by rank of the male which group they belong.
Due to change of habit of life proportions of animals have changed that was
resulted in their behaviour. More graceful constitution has made impossible
struggle by ram heads impacts characteristic for ancestors. Males sort out
their relations striking each other lateral impacts by head. Crushing blows
characteristic for rams were transformed at these animals to demonstration
of horns. The sylvammon male shows itself to the contender having bent head
down. Thus horns are lifted upwards that visually exaggerates his size. At
this time mane on neck of the male is fluffed. Males rear on back legs and
can make some steps in such position for strengthening of the impression created
at the opponent.
At any season in sylvammon herds it is possible to see cubs. Regularity in
breeding at this species is not present but most part of cubs is born in spring
(in October – November in Southern hemisphere). One cub usually is born though
about a quarter of all females eans twins. Cub is born well advanced as at
all hoofed mammals. At one hour after birth it already can walk and at the
second day of life does not lag behind herd.
Great goatlope (Gravicaper magnus)
Order: Artiodactyls (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: Zinj Land, savannas and light forests at the west of subcontinent.
In human epoch ungulates of bovid family had suffered the great damage. Many
of their species had been objects of hunting, and practically all wild species
had suffered from destruction of inhabitancy and the expansion of territories
occupied for agriculture. Subsequently the significant part of these grounds
had undergone to desertification, and began even less suitable for life of
ungulates. In Neocene new groups of running herbivorous mammals had appeared,
and the variety of ruminant artiodactyls had reduced. In human epoch herds
of various wild ruminants were replaced by monospecific herds of domestic cattle
which had become extinct after people in the majority of places. But representatives
of primitive breeds of domestic ruminants became ancestors of some species
of Neocene ungulates. At the East-African subcontinent (Zinj Land) from feral
domestic goats many kinds of goatlopes had descended; these ones became dominant
herbivores of subcontinent alongside with horned suids of Porceratidae family.
In savannas at the west of Zinj Land small herds of very large ungulates similar
by a constitution to domestic humped ox or eland antelope of human epoch graze.
It is the most massive representative of goatlopes, the great goatlope. Its
genus name “Gravicaper” means “heavy he-goat”, and emphasizes features of its
anatomy. Evolution of goatlopes directed to increasing of body size is limited
by presence of large horned suids at Zinj Land. In Neocene among suids a plenty
of massive running forms occupying ecological niches of rhinoceroses, tapirs
and bulls had appeared. Therefore great goatlope is an only exception among
swift-footed gracile relatives.
Great goatlope is an ecological analogue of elands of Holocene. Growth of this
animal at a shoulder is about 160 cm, and weight is up to 800 kgs. Legs are
rather short, and hoofs are wide. Front and rear legs have approximately equal
length, therefore back of animal is horizontal. Great goatlope runs rather
slowly and prefers to live on plains with firm ground.
The large animal in conditions of a hot climate faces with a problem of heat
emitting. At great goatlope on the neck “hanger” develops – wide plica of skin
stretched from chin up to back edge of thorax. This skin is penetrated with
blood vessels, and in day heat they dilate, giving to air surplus of heat.
Skin of great goatlope is covered with very short light grey wool; from apart
its wool seems velvety. At young animals wool is rather long and darker, rather
than at adults. At adult animals long black hair growing on the bottom edge
of “hanger” form a kind of “beard” from chin up to stomach. It is especially
well advanced at males and helps to estimate force and physical condition of
the male in courtship season. On long tail there is a switch of black hair.
At great goatlope both females and males are horned. Horns of these animals
are short and have wide bases almost adjoining on forehead. Horns are lyre-shaped,
and the annual gain adds new relief ring in their basis. During courtship duels
males fall on knees of forward legs and combat, striking impacts by head and
neck. Rut at great goatlope takes place at the beginning of dry season.
Great goatlope eats lower parts of grasses and undersized plants, therefore
it prefers to graze near to other species which eat top parts of plants. This
species does not compete to bikifaru, the representative of horned pigs, because
it occupies various habitats, preferring dry ones: savannas overgrown with
grass and flat hillsides with small amount of bush.
Once a year, in the beginning of rainseason, the female gives rise to one large,
well advanced calf. One hour after birth it already can walk, and at the second
day of life it runs on a level with adult animals. Sexual maturity at these
animals comes at the age of 3 years.
Donkey goatlope (Hippocapra asellina)
Order: Artiodactyls (Artiodactyla)
Family: Bovids (Bovidae)
Habitat: Zinj Land, savannas and woodlands at the west of subcontinent.
Goatlopes evolved at Zinj Land are descendants of the common ancestor, domestic
goat (Capra domestica). Till the evolution history, occupying various habitats,
these animals changed, adapting to new conditions. It allowed them to avoid
competition and to use resources of an environment more effectively.
The highest species of goatlopes is donkey goatlope living in dry regions
at the west of subcontinent. Its growth at a shoulder reaches 170 centimeters,
and animal can browse branches at height more than 2 meters. This is quickly
running animal of gracile constitution keeping in large herds.
Donkey goatlope is easily distinguishing due to the specific colouring similar
to colouring of Grevy’s zebra (Equus grevyi) lived in human era. But the
colouring of this species is less contrast: on body, shoulders, hips and
neck on light
greyish-beige background thin and dense vertical brown strips stretch. At
old animal strips on back part of body may turn pale. At young growth colouring
is more contrast, and calves of this species seem darker. The bottom part
of
legs at animals of any age is single-color greyish-white. For recognition
of congeners donkey goatlope has dark marks on head: wide black strip at
the edge
of ears and black top part of head.
At this species horns are present at animals of both genders. They are lengthened,
twisted as at koodoo antelope, and have smooth surface. At males horns are
larger, rather than at females.
Tail of donkey goatlope is similar to horse’s one – it is short and covered
with long white hair.
Distinctive feature of this species of goatlopes is long ears. This is an
adaptation helping to be cooled. At donkey goatlope the special form of social
behaviour
is developed: as a sign of attachment to each other animals lick each other’s
ears; the female also behaves so in relation to the calf. This ritual has
practical value: saliva evaporates, improving heat emitting. Besides on cheeks
of animal
wide lobe-like hangers. If the animal becomes hot, blood inflows to them,
and they increased in size, also emitting heat in air.
Due to high growth donkey goatlope does not compete to other species of ruminants
of Zinj Land. It eats top parts of graminoids and foliage of bushes, and
can browse bottom branches of low trees. Therefore more often it lives at
edges
of small forests and thickets of bushes. This species keeps in herds including
both males and females. Such herds may total some tens animals of various
ages. Out of courtship season males are tolerant relatively to each other.
In courtship
season, which begins at the end of rainseason, they start to behave aggressively
to each other, and the herd is separated into set of harems, each of which
is jealously protected by male. At this time males start to emit very strong
“goat” smell. Males battle against each other, having widely moved apart
front legs and having lowered heads. The winner male drives defeated male
away from
harem, uttering throat bleating, and brings its females in harem. At this
moment it starts to smell especially strongly. Male of donkey goatlope “marks”
females
by original mode – he urinates plentifully on the ground, rolls in dirt,
rubring urine in wool, and then rubs against females, leaving smell of urine
on them.
During courtship season male repeats this procedure each day.
At the beginning of the next rain season the female gives birth to one well
advanced calf. In first some minutes after birth it already rises on legs
and tries to walk. At the age of three months vegetative food contains more
than
half of its diet. The female, in addition to milk, feeds calf with belched
semidigested grass, thus supplying it with unicellular organisms necessary
for life, which live in stomach at all ruminants.
Young females become sexually mature at the second year of life, and males
at the third year. Life expectancy of these animals in nature does not exceed
20 years.
Duikapi
(Giraffotragus altus)
Order: Even-toed ungulates (Artiodactyla)
Family: Bonehorns (Osseocorni seu Parabovidae)
Habitat: savannas and light forests of Southern Africa.
After the ice age of boundary between Holocene and Neocene borders of some
zoogeographic areas have changed. So, after Mediterranean sea drying and Gibraltar
isthmus formation border of Holarctic zoogeographic area has moved to the south
having included the north zone of African savannas (the middle Sahara watercourse
of Niles (flowing now from the east to the west of Northern Africa and inflowing
to headstream of Niger river) serves as border between Holarctic and Ethiopian
zoogeographic areas). In Northern Africa there are descendants of Eurasian
and Near Eastern animals and small impurity of descendants of African forms.
But in Southern Africa separated from the north by continuous zone of mountains
and marshy woods stretched along the equator relicts of a Holocene have remained
and evolved - various species of even-toed hoofed mammals, descendants of different
African antelopes. They occupy different ecological niches and are not less
various than species of antelopes in early historical epoch.
Among herds of new South-African antelopes grazing in dense grass giants sedately
wandering by small groups and regaling themselves with soft leaflets of trees
inaccessible to anybody from other hoofed mammals are especially remarkable.
These giants of Africa are duikapi – descendants of one species of the African
wood duiker antelopes (Cephalophus). Though ancestors are too small, their
descendant is rather large herbivore mammal: its withers height is up to 170
cm at body length up to 2 meters. Neck of duikapi is very long (the common
growth of animal from horns up to hoofs is almost 3 meters). Colouring of short
animal’s wool is sandy-yellow with brown cross strips on legs and neck; back
is dark. Cubs of duikapi seem darker than adults: on their body strips are
narrower and denser.
These long-legged animals move with amble similarly to giraffes and camels.
In case of danger they can run gallop accelerating momentum up to 50 kilometers
per hour.
Skull of adult animals is strongly bent in middle part; large eyes and small
brain cavity tower above the lengthened obverse part. Due to high position
of eyes at these animals there is fine circular field of view and they notice
predators from afar. Similarly to giraffes and ostriches in Holocene epoch
savannas duikapi serve as though as “sentinels” for other herbivores. Large
nasal cavities indicate that at duikapi sense of smell is very keen.
On nape of duikapi the original cross wide “horn” is developed covered with
wool and doubled at top where reduced rests of true horns, two small horn knobs
are placed. Such bone ledge is used as the tournament weapon: contenders approach
to each other by chest, rest foreheads and “struggle” by heads trying to curve
neck of the opponent to force it to refuse struggle and to recede. Such fight
is the original compromise between natural habit to butt inherited from bovine
ancestors and rather fragile constitution of duikapi.
Duikapi keep in groups of 5 - 8 animals under the leading of elder female.
In such group there are some adult females and their under one year aged cubs.
Elder young males abandon mothers and young females can remain in parental
group or leave it and create own one. Males live solitarly or in groups of
2 - 3 animals. Out of breeding season they tolerantly concern to each other
but in breeding season their aggression relatively to each other sharply increases.
Duikapi are out of competition among animals of South-African savannas and
light forests. These high animals browse branches of trees and bushes inaccessible
to their neighbours. If all accessible lower branches on tree are eaten duikapi
easily can rear on back legs resting hoofs against tree trunk and reaching
up to fresh branches.
Pregnancy at this species lasts about 6 months. At the female only one cub
is born at the end of dry season. Growth of newborn cub is about one and half
meters. First time it feeds exclusively by milk but from monthly age gradually
passes to vegetative food. It stops to depend on mother at the age of half-year,
and becomes completely adult at the third year of life.
Thick-horned
thunderhorn (Brontocornus magnus)
Order: Even-toed ungulates (Artiodactyla)
Family: Thunderhorns (Brontocornidae)

Habitat: Hindustan, South-Eastern Asia, woodlands, bushes, foothills.
In human epoch representatives hoofed mammals had a hard time. They appeared
among favourite game, number of many species had strongly reduced, and some
of them had disappeared at all as a result of hunting or change of habitats.
The ice age had finished this epoch appeared one more test for them. Many species
at this time had disappeared and variety of hoofed mammals had sharply decreased.
Some orders like proboscid and sirens had disappeared completely. Therefore
in Neocene hoofed mammals are presented by not numerous isolated families.
But their evolution was not finished - survived groups of animals evolved to
descendants perfectly adapted to changed inhabitancy.
In India separated from other Eurasia by Himalayas the whole new family of
even-toed mammal had appeared. Its representatives differ in various form of
horns turned to resonators for strengthening of voice. Because of this feature
animals had been named “thunderhorns”. They descend from bovines, obviously
from any small antelopes. Cavities of bovine horns had joined frontal and nasal
cavities. At thunderhorns, their descendants, these cavities had got more complex
form and horns had turned to resonators. At different species of these hoofed
mammals resonators have various shape and voices of these animals differ appreciably.
The voice serves for recognition of species representatives in bushes at a
great distance (all thunderhorns are inhabitants of forests and bushes). One
more role of loud voice is to inform the probable contender that any territory
is occupied and by that to avoid force methods of struggle. Resonator outgrowths
are rather fragile ones, also they are unsuitable for force struggle. In this
connection the horn cover on these outgrowths at different species is at different
stations of reduction up to its full disappearance. Females at all species
of thunderhorns have fewer horns than males.
Thick-horned thunderhorn, the central representative of family, has kept more
primitive ancestral features than other species. Its horns are similar to strongly
swollen and short ram horns having kept horn cover on the top side. The voice
of the present species resembles roar of any predator.
Thick-horned thunderhorn is small deer-sized animal differing in graceful constitution.
The basic colouring of wool is light brown. Neck, back and rear legs of animal
are covered with short black cross strips forming fine cryptic colouring. On
legs black wool forms “stockings”. Tail is white, on the bottom side of neck
also lengthened white wool grows. Tail put upright means signal of danger.
This animal keeps among bushes feeding by leaves and grass.
Thick-horned thunderhorn does not form herds and keeps in “harems”: one male
and 4 - 6 females. In breeding season between males duels take place accompanying
with loud frightening roar audible for kilometer and more. The rut falls to
the beginning of dry season. Pregnancy lasts about 5 months. As a rule, female
gives rise to one calf colored similarly to adults but having longer strips.
At the age of one year young animals leave parental herd and form their own
herds. Sexual maturity comes at three-year age (at females it happens some
months earlier, than at males). At this time herds of young animals separate
to harems. Life expectancy in nature reaches 12 - 14 years.
Forest
trumpeter (Buccinotherium magna-voce)
Order: Even-toed ungulates (Artiodactyla)
Family: Thunderhorns (Brontocornidae)

Habitat: Hindustan, foothills of Himalayas, light forests and thickets of bushes.
This species is large hoofed mammals of heavy-build constitution, one of the
largest representatives of family: withers height of the adult male reaches
2 meters,
female is a little bit smaller. The skull of this species is rather long; more
than half of it the hollow resonating crest of the roundish shape amounts.
Horn covers have completely disappeared and remained bases of horns are covered
with velvety wool and rather brightly colored. The crest is high, compressed
from sides; its edge is bordered by wide strip of black wool, and the middle
part of crest is reddish-yellow. Body of this animal is colored bluish-gray;
stomach is white. On long tail rich brush of white wool grows. Position and
character of movements of tail at this animal expresses its condition at present
moment.
The voice of forest trumpeter has determined its name: it sounds like lingering
blares.
Jaws are long and rather weak: animal eats mainly soft food - leaves of trees
and bushes. To browse it from branches on top jaw nose and upper lip have formed
similarity of short proboscis, as if at tapir.
Wood trumpeters live in forests near large rivers where giant grasses and bushes
plentifully expand. Hoofs of this animal are well adapted to walking on fenny
ground and then forest trumpeter is able to swim well.
This mammal lives by couples kept for the whole year. The male incurs protection
of territory borders marking them by heaps of manure. Also he warns the probable
contender that the territory is occupied with the help of shouts. Wood trumpeters
are especially “vociferous” in the morning. They actively shout while above
river valleys there is dense fog: probability of predator attack is less so.
Once a year, mainly at the end of dry season female gives rise to one cub.
By colouring it is similar to adult animals but on the head instead of crest
only strip of black wool is. The young animal keeps with parents almost year,
and female banishes it shortly before birth of the next cub. The animal becomes
the adult at the age of four years.
Shadhavar
(Shadhavar fabulous)
Order: Even-toed ungulates (Artiodactyla)
Family: Thunderhorns (Brontocornidae)

Habitat: Hindustan, Southwest Asia up to the Persian ridge, plains and light
forests.
In Oriental (Persian) legends shadhavar is the fantastic creature : the antelope-like
one with unique hollow horn. According the legend when the wind blew, the horn
of this antelope sounded attractively to other animals. When curious animals
approached closer, shadhavar rushed on them and ate: the word of mouth attributed
to this fantastic animal furious behavior. And only the voice of the wild pigeon
could charm this animal so that hunters could catch it.
But legends have disappeared together with people, and in savannas and light
forests of Southern Asia the animal somewhat resembling the legendary creature
has appeared. Its behavior is not so furious, as at the fantastic prototype,
but some other properties rather approach this hoofed mammals from thunderhorn
family with fictious creature. The horn of this animal is especially remarkable:
bases of pair horns are pulled together and have grown together to the single
structure resembling the crest of dinosaur Parasaurolophus. Only this hollow
horn is straight, also continuing a line of animal head. The length of horn
exceeds length of other skull in one and half time. The corneous cover on this
structure has disappeared completely; "horn" is covered with short
thin wool. It is not more used for fight, its purpose is to be the resonator
and partly to carry out alarm function. The tip of horn is covered with black
wool: it is a mark for recognition of neighbours. During ritual combats males
bend head down so that the horn sticks upwards as it is possible. This way
they determine the strongest one without fight. The horn of the female is shorter,
and the black tip on it is not so big. Due to horn the male can utter lingering
trumpet shout of high tone. The voice of the female is more silent, than at
the male.
By other features shadhavar resembles other artiodactyls. It is herd animal
by proportions similar to fallow deer. It lives in bush thickets in foothills
and valleys, in summer rising highly in mountains. It keeps in herds by 10
- 20 ones. The basic background of body colouring is yellowish - grey, on wool
there are longitudinal black stripes, lasting along all body from shoulders
up to hips. These stripes help animals to disappear among bushes and do not
allow predators to distinguish separate animals in herd (colouring of zebras
works by similar mode). The animal can jump high.
It eats mainly leaves of bushes and soft grasses.
The coupling season falls to the end of summer. At this time males establish
hierarchy and gather harem from 5 - 6 females, protecting them from other males.
After coupling season the hierarchy is almost erased; and recent contenders
do not pay attention against each other. Once a year, in the beginning of spring
the female gives rise to two calves. They have no characteristic horn, and
their voices sounds like lamb bleating. Calves are covered with small spots
which merge in longitudinal strips later. Approximately to the age of one year
at them the horn starts to form. At the female it reaches full development
to the age of three years, at the male – up to the fifth year of life. Sexual
maturity comes in three years, but males really can participate in breeding
from five - six years.
Desert
whistlehorn (Stridocornis aridophilus)
Order: Artiodactyls (Artiodactyla)
Family: Thunderhorns (Brontocornidae)
Habitat: semi-deserts and deserts of Near East and Northern Africa.
In human epoch hoofed mammals have strongly suffered from people activity.
Destruction of their habitats and active hunting had resulted to that only
few of them have survived up to Neocene. Indirectly reduction of number of
hoofed mammals had resulted to that other groups of animals have started to
master their ecological niches. Nevertheless, evolution of “true” hoofed mammals
in Neocene has proceeded, and they even had formed new groups which had not
existed in human epoch. One of such groups is a family of thunderhorns. They
are mainly inhabitants of open spaces and light forests of Southern Asia and
Northern Africa. One of representatives of family, the desert whistlehorn,
has mastered life in rather difficult conditions: it lives in droughty district
bordering from south and east coasts of Mediterranean salt swamps.
This animal of graceful addition looks similar to gazelle. The desert whistlehorn
is medium-sized animal: it weighs only about 60 - 70 kg, its withers height
is about 1 meter. Its legs are long and thin, with rather small hoofs. The
desert whistlehorn can run very quickly, accelerating on a short distance to
speed over 80 kms per hour. Because of small hoofs it avoids viscous ground:
sandy sites of desert and boggy saline soils. In a dry season these animals
meet on clay sites of desert, but in short rain season they leave in territory
with dense and drier ground.
Body of the desert whistlehorn is one-color, sandy-yellow; wool on back is
lighter up to almost white. Tail is long, with hairy brush of light hair on
the tip. Characteristic feature of representatives of family is resonator chambers
of various shapes on the head. At the whistlehorn they look like the vertical
semicircular “crest” covered with black cross strips on light background. As
at all thunderhorns, this outgrowth is formed by cavities of former horns,
and serves for strengthening of sounds uttered by animals. However, the outgrowth
is not only the resonator: the bone forming it is very thin also is penetrated
with plenty of blood vessels. This adaptation also serves for cooling, effectively
radiating surplus of heat. Of course, animals with such structure on head are
not able to butt, therefore relations of hierarchy in herd are established
mainly with the help of loud voices, pushes by shoulders and impacts by hoofs.
But at whistlehorns the special form of expression of submission has appeared:
weaker animal licks crest to stronger one. The sense of this behavior is, that
evaporating saliva helps the licked animal to cool. This behavior became an
element of courtship ritual of animals: the male “licks” a crest of the female
like this, but only stronger ritualized, slightly touching it by tongue.
The desert whistlehorn lives in hot and droughty district, and eats various
plants, including bitter and even poisonous for other animals ones. It can
not drink for a long time, receiving a metabolic water from the eaten plants.
This animal keeps in big herds, in which hierarchical relations are established
only in pairing season. At this time males start to utter loud sounds: long
and also shrilly whistle through nostrils, blowing air through an outgrowth
on a head (of course, it is connected with nasal ducts). Showing the resonating
outgrowth to females, the male bends head back, showing it on the background
of light wool of back.
In epoch when the climate became drier, the part of population of whistlehorns
dwelt in Northern Africa has passed the Mediterranean bogs by way of isthmus
connecting territories of Tunis and Sicily. In territory of Southern Europe
they had turned to the special species – Etrurian whistlehorn (Stridocornis
etrurius). The European species differs from African one in larger sizes, dark
colouring (wool on sides and back is monotonous red), and propensity to life
in foothills. Etrurian whistlehorn runs not so quickly, but dexterously swarms
up rocks. It has stronger constitution, and its forward legs are appreciably
shorter then back ones: it is a feature of adaptation to inhabiting on mountain
slopes. Head of this animal is narrower, but higher, with short and high resonating
crest. Courtship cry of this animal also differs from voice of desert whistlehorn:
it sounds like abrupt loud sounds.
Mountain
camelope (Leptocamelus montanophylus)
Order: Tylopods (Tylopoda)
Family: Camels (Camelidae)
Picture
by Rafael Silva do Nascimento, Brazil
Habitat: mountain areas of Australia.
In Australia camels never were found in wild and they could not get to this
isolated continent naturally. Woods and sea passages reliably barred them the
way to the “lost world” of Australia. Only due to one species of primates,
Homo sapiens had become extinct at the end of Holocene they could get to this
continent as domestic animals. After human disappearance their ancestors, dromedaries
(Camelus dromedarius), have become wild and have given rise to several species
of herbivores competing with local marsupial animals. Among camels analogues
of flatland giraffes and antelopes, and also mountain goats have appeared.
The mountain camelope is strong build relative of swift-footed camelope Leptocamelus
gracilis, graceful runner of Australian plains. This species differ from plain
neighbour in shorter legs and neck, and also in larger narrow head. Besides
mountain camelope is covered with dense wool that helps to endure cold night
of Australian Alps. It is rather medium-sized species - its weathers height
is up to 1,2 m, body length is about one and half meters. Back of mountain
camelope is inclined forward because back legs of animal are appreciably longer
than forward ones – so it is more conveniently to graze on mountain slope.
Wool of animal is colored grey with brownish shade; on ridge there is longitudinal
black strip which at males is much wider and longer than at females. The hump
at these camels is expressed poorly - in middle of back small swelling is appreciable.
On chest and basis of neck at animals of both sexes mane of long hairs develops.
On head at males very long hairs grow a rich bun. To time of rut a liquid from
specific gland located on nape impregnates them – hair help to strengthen smell
and to keep it for a long time. At females such hairs are not present; head
is covered with short wool. Legs are ended as at all representatives of the
order by two hoof-like claws and special skin “pad”; the weight of body falls
to end toe phalanxes. For the best cohesion with stone surface leg soles are
covered with strongly cornificate skin.
Mountain camelopes live on western slopes of Australian Alps in conditions
of cool and rather dry climate. These animals prefer to live in forestless
districts not coming to thickets of trees occasionally growing in valleys.
They eat grass at times reaching up to plants on very abrupt slopes. Due to
features of legs these animals easily jump and swarm up hillsides surviving
in places where marsupials of size comparable to them can not live. Teeth of
mountain camelope have the special structure: incisors are strongly jutting
out forward as if a nipper, and molars grow all life: it is an adaptation to
grazing of food plentifully “flavoured” with sand and dust strongly erasing
teeth.
These animals live in herds of 10 – 15 ones under the rule of large male leader.
He constantly supports the authority in herd with the help of might demonstrations
and loud roar. In herd the significant part is presented by mature females
and their cubs. At mountain camelopes one cub alternate years is born as a
rule. It depends on mother for a long time studying at it to move in mountains
and to search for food. Usually it becomes completely independent at the third
year of life leaving mother at birth at her of the next cub. The leader of
herd does not support youngsters in herd as among them his probable future
contenders can grow. When young animal leaves from under parent trusteeship,
it has a lot from herd leader forcing young growth to submission. Because sexual
maturity at young females comes only at the fourth year they do not represent
interest for the main male and also are driven out. Young animals form small
herds either mixed or including only young males. At this time the part of
young growth perishes from various predators. Such herds of young growth live
in boundary territories between areas of breeding herds and occasionally between
old and young animals skirmishes happen. When young females become sexual matured,
between young males fights for “harem” possession begin. Contenders rear, collide
by chests, strike impacts by forward legs in sides and chest of contender and
also bite strongly. Thus fighting males loudly roar and sometimes such loud
demonstration of ardour serves bad service for them: having heard such roar
skilled adult males hasten to these sounds and disperse self-confident young
growth adding some young females to own herd. If at the herd “sharing” place
two adult males face, fight between them happens severe and frequently results
to mutilations or fatal outcome. Sometimes young male can live many years without
“harem”, from time to time attempting to “property” of any “married” neighbour.
Sometimes such bachelor literally chases herd of the contender trying to deliver
from it one or several females.
Mountain camelopes are threatened, as a rule, only with local marsupial predators.
If there is no opportunity to rescue in flight, herd can pass to active defense
trying to strike predator by forward legs or to bite.
Life expectancy of mountain camelopes can reach 40 years.
Caballocamelus, horse camel (Caballocamelus velox)
Order Tylopods (Tylopoda)
Family Camels (Camelidae)
Habitat: savannas and semideserts of Meganesia.
Up to human arrival it was no large running mammal in Australia. Birds and
kangaroo were their ecological analogues. The occurrence of camels (Camelus
dromedarius) introduced by people had changed the course of evolution of continent
inhabitants. Marsupials could not evolve to four-footed quickly running animal,
replacing hoofed mammals only from the point of view of ecology. Despite of
the competition to camels, they had survived and successfully evolved in Neocenic
Meganesia. They avoid a competition to descendants of camels due to feeding
by different kinds of forage, and also due to effective digestion of accessible,
but poorly nutritious food. As a result descendants of camels had occupied
only a part of ecological niches in plains of Australian part of Meganesia.
Among them there are camelopes of gracile constitution, and high giraffamels having no competitors in bottom and middle levels of tree tops. But these animals
live in districts where there is though any wood vegetation. In plains of Central
and Western Meganesia, to the south of the zone of forests and woodlands, another
descendant of camels, caballocamelus, the gregarious mammal similar to the
horse, lives.
Caballocamelus is a large animal of robust constitution: its withers height
is up to 160 cm, and the general length of body is about 3 m. At this animal
there is a large head on rather short and strong neck. Jaws are short and have
constantly growing teeth, adapted for grinding of rigid graminoids and bush
branches. The muzzle is extended to short almost immovable proboscis serving
for protection against dust and humidifying of inhaled air.
The hump is small; it is stretched along the back up to waist as the low wide
platen. The tail is turned to “fly-brush” having hairy brush on the tip.
The wool of caballocamelus is very short and velvety. On the bottom part of
neck at adult animals a lot of lengthened hair similar to beard grows. Colouring
of body is yellowish brown; on shoulders and hips there is a small amount of
faltering dark strips. On legs of animal there are white “stockings”. At adult
animals head is darker, rather than body; at male forehead and nose bridge
have coffee-brown color. At young growth the head has the same color as the
body, and on legs there is lack of “stockings”.
The foot of typical camel is adapted to rather slow movement on soft ground:
toes are widely separated, sheafs between them are lost, and the support falls
to some distal phalanxes. At caballocamelus the structure of foot has changed
according the adaptation to fast running. Toes of this animal are connected
by sinews to each other along almost all length. At young animal sinews are
elastic, but to the approach of maturity they ossify. Distal toe phalanxes
are thick and covered with “stockings” of thick skin. The characteristic for
camels subungius, at which the leg bases, is very small.
This animal lives in dry districts with firm ground, lack of wood vegetation.
Caballocamelus quickly runs, accelerating momentum about 50 kms per hour.
Its pace at short distances is gallop, at long one it moves by an amble (it
is
a characteristic
pace of typical camels). This species lives in herds numbering 20 – 40 animals.
Caballocamelus eats rigid grasses and can even gnaw prickly branches of bushes.
Its three-chambered stomach (inherited from the dromedary, its ancestor) helps
to digest such food. Lips covered with dense skin help to feed on prickly plants.
In rain season animals can not drink till many days, being content with moisture
received from plants. But in dry season these animals visit reservoirs approximately
once in three days. The leader of herd knows well all reservoirs in places
of inhabiting of herd, and each time animals visit new reservoir for watering.
The herd includes from 1 up to 3 harems leading by dominant males. Each male
owns 5 – 10 females; their posterity stays in herd up to the maturity (young
males are expelled from the herd a little bit earlier, rather than females).
Young females are more often quietly poured in other herds, and single males
form herds. Becoming stronger, they try to bereave any adult male of authority.
Males of this species are very aggressive in courtship season: they bite each
other and beat by forward legs. Skin of the male is thick, especially on shoulders
and in the basis of neck – it is the adaptation for intraspecific courtship
tournaments. Out of breeding season males treat to each other more tolerantly,
but at the watering place harems approach to water in sequence appropriate
to rank of the male in herd.
Pregnancy lasts about 10 months; one cub is born always. The female cares of
it for a long time, therefore the posterity at the female is born only once
in 2 years. The female does not admit neighbours to the newborn cub while it
will rise on legs, and some more hours after it. At this time the cub remembers
smell and voice of mother. It is very much adhered to mother, and does not
depart far from her for a long time. A parental instinct at this species is
very strong: in case of need all adult animals of a clan protect the cub; therefore
the survival rate of posterity at this species is rather high.
Siberian
digger (Paramarmota architector)
Order: Rodents (Rodentia)
Family: Squirrels (Sciuridae)
Habitat: Western Siberia, flat districts, river valleys.
Small animals had gone through mass extinction of the epoch at the boundary of
Holocene and Neocene much more successful, rather than large species. Their variety
at the level of families remained practically the same, as before the catastrophe;
separate genera had died out, but new ones appeared to replace them. Ecological
niches occupied by them had remained the same – these are small forest and grassland
animals, herbivores and omnivores. Among them also digging forms remain, for
example, suslic doggies from three-Rivers-Land steppes. In severe continental
climate of Siberia the separate species of digging animals settling in large
colonies, Siberian digger, also have appeared. The colonial habit of life provides
more successful survival of these animals in conditions of contrast continental
climate and severe long winter. This animal descends from one species of suslics
inhabited glacial steppes during the congelation at the boundary of Holocene
and Neocene.
From the point of view of anatomy Siberian digger represents very large suslic
– one average animal weighs up to 3 kg. Ecological analogue of Siberian digger
is the marmot – digging animal building extensive underground constructions.
The colony of Siberian diggers numbers up to 200 – 300 animals, and they construct
under the ground a huge settlement which part is placed at the depth up to 4
– 5 meters, inaccessible for predators.
Settlement of these rodents represents the circuit of underground tunnels supplied
with drainage “shafts” and drainage burrows which open in steep riverbank and
can stretch to tens meters from edge of colony. It is necessary for protection
of colony in spring from thawed snow. Besides exits to the surface are made as
high (up to two meters) earthen hills with an entrance at the top. At snow thawing
in them only a little of ice-cold water will get. From these hills the good view
opens, therefore in summer at tops of hills “sentries” – males from among defenders
of a colony –are constantly on duty.
Except for earthen hills – entrances to the colony, presence of colony is marked
by the bushes and young trees damaged by rodents. Siberian diggers influence
a landscape as essentially, as large herbivores like obda or shurga: within the
limits of their colonies trees survive hardly (they are simply chewed at the
sprout stage). But above colonies of diggers graminoids and other plants, bearing
drier ground (presence of drainage system in colonies has an effect) and constant
“shear” of them by these rodents, prosper. Forager individuals from colony also
make sorties for the forage in thickets of bushes and forest.
Siberian diggers are animals having well expressed caste system and obvious intraspecific
polymorphism. Like Neocenic castle rabbits from New Zealand or naked mole rats
(Heterocephalus glaber) from Africa of Holocene epoch, these animals are divided
to castes. The caste system is not fixed genetically, and depends on age and
position of animal in hierarchy of colony. At these rodents there is large head
with very short ears, able to wrap when the animal moves in hole. The trunk of
Siberian digger is covered with rich velvety fur similar a little to mole fur:
hairs grow vertically, and do not prevent movement in hole forward or back. The
fur of animal is colored brown with small amount of white spots on back and shoulders.
On head of adult males there is an area of white fur on sides of head and around
of eyes. Depending on position of animal in hierarchy colouring of fur may vary
strongly.
Paws of Siberian digger are short, with wide feet, having edge of rigid hairs
–it is convenient to rake away the ground by such paws during the digging. Tail
is short, with white tip. Tail lifted upwards is an alarm signal, well appreciable
even from among grass.
The caste system of Siberian diggers includes some categories of animals:
1) “King” and “queen” – breeding pair of large fertile animals. But the amount
of these animals is not limited, as at insects. In large colony occurrence of
several “queens”, which are fertilized by one or two “kings”, is possible. If
two “kings” meet in tunnels of colony, between them severe combat is possible,
but usually “king” does not leave from the centre of colony. From these animals
all posterity of colony descends.
“Queen” represents the individual which almost continuously gives rise or brings
up young growth. It lives in special chamber; it and its posterity are looked
after by some females from the number of “nurses”. “Queen” is very large – it
is approximately twice larger than the adult forager female. At it milk glands
are strongly advanced, and it almost continuously produces milk (except for second
half of pregnancy). Because of it “queen” requires a plenty of juicy forages,
and “nurses” additionally give it to drink, dragging water from drainage “shafts”
or from the nearest reservoir in cheek pouches. Fertility of “queen” is about
15 naked, blind and helpless cubs in one litter. In some days after cub birth
it is ready to become pregnant again. For one year “queen” makes up to five litters.
“Queen” can not look after itself and posterity independently; it is only able
to creep hardly in the chamber. If in spring thawed snow fills in chambers where
“queens” live, they perish, being not able to escape.
“King” is the large fertile male with normal proportions of body, able to serve
itself independently. It freely moves in colony, independently eats in “pantries”
and searches for “queens” ready to fertilisation. At this male sexual attributes
in colouring of wool are obviously expressed: on head around of eyes and on cheeks
areas of white wool develop. Also “king” has repugnatorial glands: it has a specific
smell which influences other males and suppresses sexual behaviour at them.
2) “Soldiers” – aggressive adult males. They are in condition of constant stress
because of presence of “king” in colony. The stress causes in them formation
of plenty of adrenaline, and hormonal changes lead to irreversible changes in
their appearance. Incisors and jaws of “soldiers” become large; therefore the
head of such individuals looks disproportionately big relatively to the body.
Sexual function at them is suppressed from the childhood, especially if they
developed in conditions of density in the centre of colony. These animals can
hardly eat independently because of hypertrophied jaws; more often young individuals
from “nurses” caste feed and clean them. But at an attack on colony they leave
on surface the first, and furiously attack any animal had disturbed them, even
such large one, as obda or shurga. At the edges of colony where the stress is
less, features of “soldier” at males are less expressed, and among them fertile
individuals, from whom new “kings” may grow, frequently meet.
Colouring of “soldier” is actually warning: at them there is almost completely
white head with dark marks on forehead and nape, and white spots on shoulders
are very big. Such colouring is well remembered by animals which had undergone
to attack of “soldiers” of colony, and forces them to keep farther from these
animals.
3) “Foragers” and “nurses” – females of various ages. Adult and old females are
foragers. They constantly gather forages and drag them in holes serving as “pantries”.
These animals are able to feed and look after themselves independently. Young
females have function of “nurses” – they look after “queens” and “soldiers”,
clean and feed them. At them the parent instinct is early expressed, but it equally
quickly fades. If in the short period of maturing the female was not fertilized,
it becomes the forager. Young “nurses” have opportunity to become “queens”, if
they will be fertilized within first three months of life. At pregnancy there
is a hormonal reorganization of the organism, and young female develops to “queen”
occupied with reproduction.
If the colony becomes too large, in it “diarchy” can arise: at the edge of colony
the young male (from among the individuals less subject to stress) and the female
ready to breeding from the number of “nurses” can create new breeding family.
If founders of colony perish, their place is occupied quickly by individuals
from among young ones. Frequently young males from other colonies come in colony.
They lodge at edges, and also can become founders of new “dynasty”.
Siberian diggers eat exclusively vegetative food. Everything, that can be stored
within winter, is reserved in special chambers serving as “pantries”. These are
tubers and rhizomes of plants, and also seeds of graminoids. Daily food of animals
in summer includes young branches of bushes, greenery and roots; the autumn adds
to them fruits of various plants. Foragers of this species carry food in cheek
pouches. In them there are no salivary glands, therefore the transporting forage
stays dry and does not spoil in winter.
In winter activity of colony is reduced: animals spend the most part of day in
dream. When bad frosts come, all colony runs to true hibernation, at which the
body temperature of animals is strongly reduced, and vital processes are inhibited.
At this time even development of embryos at breeding females stops. Due to this
feature consumption of forages does not increase, and the colony successfully
exists up to the end of spring when there is an opportunity to pass to fresh
forage. When snow thaws, flooding of the part of colony is possible, but the
majority of animals usually survives. When all difficulties are left behind,
and in nature there is enough of fresh vitamin forage, in colony the burst of
birth rate takes place. The care of posterity raises survival rate of young growth,
and damage, caused to the colony by predators and weather, is restored.
Life expectancy of foragers and “soldiers” seldom exceeds three years, but “kings”
and “queens” can live till ten years and more.
Farmer
hamster (Agrocricetus agricola)
Order: Rodents (Rodentia)
Family: Hamsters (Cricetidae)
Habitat: steppes of Southern Europe (Three-Rivers-Land) to the east up to Southern
Ural and Central Asia.
Rodents are the most numerous global widespread order of mammals. Their species
increased variety till all Cenozoic era and had got advantage in Holocene.
Human economic activity in many cases promoted prosperity of rodents. People
superseded and destroyed wild hoofed mammals, ploughed steppes and cultivated
large plantations of graminoids. People had also exterminated many feathered
and four-footed predators, and it had favoured to breeding of separate species
of rodents. They grasped new habitats freely, getting of advantage from human
activity.
In Neocene extensive spaces in Southern Europe, Middle and Central Asia had
turned to grassy plains in which local herbivores graze. Their typical representatives
are steppe species of harelopes and massive porcippula similar
to heavy-build pony. They “cut” steppe grasses by teeth, but do it not everywhere.
Here and there in steppe sites are located, where the grass is much denser
and higher rather than usually. Graminoids in these places differ in rather
massive ears rocking in the wind – it is a separate species, the descendant
of feral wheat many MY ago cultivated by people. These thickets differ in strict
monotony - among them plants of other species are practically absent. Obviously,
these plants were specially selected and carefully protected - such sites are
precisely separated from other grasses. Local herbivores for any reasons avoid
eating this grass which looks much more appetizing, than everywhere.
The puzzle is disclosed very simply when among high grasses there are small
stumpy mammals with rich longitudinal – striped fur of gray-brown color. It
wanders in thickets of high grasses, from time to time stopping and breaking
off stems lasting among grasses. This is one decision of some puzzles at once:
the rodent really looks after this graminoids. Actually, these thickets represent
its extensive fodder site. And due to it the animal is named farmer hamster.
This animal is a little bit smaller, rather than common hamster (Cricetus cricetus),
the usual inhabitant of Eurasian steppes of Holocene epoch. Probably, it is
its descendant though it is possible, that its ancestor could be one of small
species of Central-Asian hamsters. Colouring of farmer hamster combines two
patterns. The top side of body of this animal is cryptically colored: on gray-brown
background narrow black strips stretch. Having dropped to the ground, this
animal becomes completely imperceptible from afar for feathered and four-footed
predators. Coloring of stomach strongly differs from coloring of the back:
on stomach and cheeks there are wide cross black strips on white background.
This coloring becomes appreciable from apart when this small mammal rears to
eat something or simply to look round. At this animal there is constitution
typical for hamsters: large head, short thick trunk and very short tail, hardly
appreciable among wool.
This mammal differs in unique strategy of survival. It not simply gathers seeds
of plants which serve as food to it, and specially creates conditions for their
growth. It carefully keeps up a site of steppe around of the hole and in summer
selectively eats plants which do not belong to graminoids. As the result at
the “allotment” occupied by this hamster amounting about one hundred square
meters graminoids grow well, providing the hamster by grain for winter. More
often at sites of farmer hamster the hamster
wheat grows – it is the special genus of graminoids again “cultivated”
by the hamster after people and actually entered symbiosis relations. In summer
seeds of graminoids have not ripened yet, and animal eats grass – weeds growing
at its site. Also it eats various insects, involuntarily relieving fodder graminoids
from wreckers. In the autumn when seeds of grasses ripen, farmer hamster gathers
“crop”: it chooses the ripest ears and carefully husks them, scattering dust
on site and carrying off in burrow well cleared grain. Gradually at the territory
“cultivated” for a long time graminoids start to grow especially well, and
obviously prevail of other plants.
Farmer hamster also fertilizes ground with dung – it does not have specially
removed “toilet”, and the part of mineral substances comes back in ground with
its dung. Only in winter it leaves dung in special chamber of hole, and throws
it out outside in common with used litter in spring - it additionally fertilizes
ground in the territory this way.
Well-groomed sites of these mammals are favourite object of attention from
the part of other grain and grass lovers. At sites of farmer hamster in autumn
birds feed, and large herbivores occasionally visit them. But the farmer hamster
can render their repulse: despite of small growth, it is very much martial
creature. It frightens off smaller pilferers by loud squeals and high jumps.
Thus small mammal strongly puffs cheek pouches to seem larger. Such reception
does not work against the large herbivorous animals: they are strong enough
simply to crush this animal by hoof. But the hamster uses one more effective
means of protection against them: it lies down on back, showing to the uninvited
visitor contrastly colored belly, and shrilly squeals. If threat has not worked,
it sprinkles to the stranger stinky substances from strongly advanced anal
glands similarly to skunk.
The specific habit of life had transformed this animal to even greater “homebody”,
rather than hamsters of Holocene epoch. But it does not avoid the society of
neighbours, and prefers to settle in rarefied colonies. In total “allotments”
of different individuals of this species gather to the colony rather closely
to each other - their borders are divided with some meters. In the center of
colony there are “allotments” of strongest individuals – they are richest and
better protected from herbivores. But in the centre of colony lack of place
is sharply felt, and for each superfluous plot of land a real battle takes
place. Individuals at edges of colonies feel first that they are not unique
local herbivores. But it is possible for them to expand their “allotment”,
having “cultivated” more parts of steppe. The certain balance and order in
a colony is reached so.
Outside of breeding season each individual diligently marks borders of the
territory, sprinkling on grass secretions of anal glands. Only being ready
for pairing the female can leave the site to meet the male – in any other time
it would meet her with bites of sharp incisors. In courtship season the fodder
site is an original secondary sexual attribute: the female chooses the male
for pairing looking the size and “cultivating” degree of its allotments. After
pairing female returns to the site and rears posterity itself. Within one summer
the female rears two – three hatches of 4 – 6 cubs. The most part of young
animals settled from colony can not find suitable site, and perishes in teeth
and claws of local predators. To the autumn survived young hamsters find site
for life, and gather in hole the first stock for winter – basically seeds of
wild-growing grasses. In autumn the farmer hamster fattens up intensively and
grows fat strongly: fat may make up to half of weight of animal.
In winter farmer hamster falls into deep hibernation: the temperature of its
body is reduced practically up to an ambient temperature, and pulse decreases
up to several beatings per minute. Some times for winter animal wakes up to
eat some of seeds from the stocks. It is awake till some hours, yet will empty
intestines completely, and then falls runs into hibernation again.
Though farmer hamsters are convinced and bilious egoists, all of them keep
the contact with each other: between their holes there are burrows connecting
them to whole system. To tell the truth, more often other steppe animals dig
these courses, but hamsters willingly use them. In such tunnels animals leave
from chasing of predators. Being on the ground surface, animals “communicate”
with each other with the help of whistling, as suslics. In such way they warn
each other of occurrence of predator – usually steppe
eagleraven or predatory mammal zibetonyx hunt
them.
The biggest complexity in life of farmer hamster is derivated by its habit
of life, to be more exact – by the way of “agriculture”. At their sites only
the unique species of graminoids frequently grow, therefore animals endure
heavy time when cultivated plants are attacked with illnesses and parasites.
On “hamster wheat” the ergot fungus grows often. But small ergot infection
of graminoids does not harm to animal – when this fungus appears, hamster specially
searches for such ears, chews fungus off from them eats. Ergot poison renders
intoxicating effect to these rodents – in the beginning animal becomes easily
excitable and aggressive, then gets into the hole and sleeps for a long time.
The activity of farmer hamster constrains the distribution of ergot, but at
mass disease of graminoids they can not cope with it, and the part of crop
perishes. After flare up of plant desease in colonies of farmer hamsters the
famine comes, and number of populations may decrease in tens times. And in
such situation the part of colony members simply leaves on new places where
new “plantations” appear in some years.
This species of animals is discovered by Momus, the forum member.
Whistling
hamster (Buccacricetus stridor)
Order: Rodents (Rodentia)
Family: Hamsters (Cricetidae)
Habitat: steppes of Tree-Rivers-Land, thickets of grasses.
Some species of rodents had received great advantages in survival in human
epoch. Expansion of agricultural areas had allowed them to settle widely; the
cultivating of cereals had provided constant and uninterrupted source of food,
and human presence and its activity had strongly reduced number of predators
in habitats of rodents. Disappearance of mankind has restored natural balance
of environment during several hundreds thousand years – the extensive areas
of monocultures had disappeared, predators had appeared, and gradual formation
of new ecosystems instead of ones destroyed by people began. Evolution had
gone in the natural way again, and at new species of animals new adaptations
for survival began to develop.
In extensive steppes of Tree-Rivers-Land reaching along northern and eastern
coasts of Fourseas, rodents are prevailing group of mammals. Quickly evolving,
they had formed many species, differing by requirements to inhabitancy. Representatives
of hamster family differ from other rodents by ability to gather the large
stock of forage for winter. One of their species, farmer
hamster, even is engaged in original “cultivating” of one
species of graminoids on its fodder area. But other hamsters lead more
traditional way of life.
Many predators live in steppes. One of widely widespread species is zibetonyx,
cursorial species of viverrids, the most northern species of this family. It
represents danger to many small rodents, but one of the large hamsters, living
in steppe, is not afraid of its attack: it is armed with sharp incisors and
is able to protect itself. This species is whistling hamster.
Whistling hamster is rather large rodent: the length of its body is about 40
centimeters, and weight reaches 2 kgs. By appearance it is similar to other
hamsters: it is short-legged and massive one with large head. Its wool on back,
nape and top part of neck has camouflage color – golden-yellow with darker
faltering longitudinal strips. The bottom part of body sharply differs by colouring:
stomach and forward part of head are white with sparse black spots.
The most remarkable feature of whistling hamster is its cheek pouches. Usually
they are used for dragging stocks of forage to the hole, but at the attack
they are used in completely unusual mode. The aggressive animal rears on hind
legs and starts to puff cheek pouches. Their skin is very extensable, and due
to it this rodent seems larger and more impressive. And then remembered colouring
of wool is expressed in full degree – on skin covering cheek pouches - cross-striped
pattern becomes visible. Skin under wool is also pigmented at the places where
black hairs grow; therefore this pattern stays appreciable, even when cheek
pouches are puffed completely. This is the first warning to predator. Usually
it is enough to the skilled predator to stop getting into touch with this rodent.
If attempts of attack proceed, hamster contracts ring muscles encircling cheek
pouches, and blows them off. Thus loud whistle is hearing. This is more powerful
prevention to predators. After that hamster rushes on the enemy and seizes
it by incisors. It happens, these animals attack even porcippulas –
large and harmless herbivores.
Whistling hamsters live solitarily in rather extensive territories – between
holes of separate individuals there is a distance not less than 200 meters.
Warning relatives that the territory is occupied, the owner of territory whistles
from time to time, puffing cheek pouches. Having heard its whistling, neighbours
usually also respond.
This hamster gathers seeds of graminoids in its extensive individual territory,
and to the end of an autumn it gathers in underground “pantry” up to 30 kgs
of grain. In addition to it, animal digs out and eats tubers and rhizomes of
various plants, a part from which it also drags to “pantry”. To winter this
hamster is strongly eaten off and runs into hibernation when autumn rains begin,
changed further by light frost and snow. For winter it wakes up many times
and eats a part of the stocks. In spring whistling hamster searches mushrooms
growing in steppe, and eats up the rests of winter stocks. Usually they suffice
up to growth of young grass.
In spring the courtship season begins at these rodents. Demonstration of puffed
cheek pouches and whistling are important elements of courtship ritual of whistling
hamster. Usually male comes itself to the territory of female ready to pairing
and starts to court after her, following her with blown up cheek pouches and
whistling. If near one female some males meet, they arrange a kind of “duel”,
trying to whistle louder and longer, than the competitor. If it does not help
to define superiority, combat begins.
Within one year the female gives rise to 2 litters of 3 – 4 cubs. Young animals
of the second litter winter together with the female and in common prepare
forage in autumn. The young growth of summer litter has time to find territory
suitable for life, to dig a hole and to prepare enough amount of food for successful
wintering. The next spring young animals from both litters can take part in
breeding.
The idea about existence of this species of animals was proposed by Simon, the forum member.
Hawaiian bamboo rat (Bambusiphagomys tector)
Order: Rodents (Rodentia)
Family: Murines (Muridae)
Habitat: bamboo woods of east slopes of mountains on Big Island (Hawaii).
In human epoch birds dwelt on Hawaiian Islands were a fine example of adaptive
radiation - the phenomena at which from one ancestor many descendants evolve.
They are connected by relationship, but adapted to different inhabitancies
and having various shape. But indigenous species of Hawaiian fauna have irrevocably
died out, not having sustained a competition with newcomer species introduced
by people. And newcomers, in turn, roughly began to evolve, showing on the
example of the same evolutionary phenomenon.
Three species of rats have got to Hawaii on human fault: black (Rattus rattus),
brown (R. norvegicus) and indigenous Pacific rat (R. exulans). The third species
was quickly superseded by descendants of larger neighbours, and had died out
till several subsequent thousand years. But two remained species had actively
evolved, having given many various descendants distinguished by appearance,
behavior and habit of life. Descendants of brown rats became mainly ground
animals, and black rat – wood inhabitants.
A few species of mammals live in bamboo woods. In this place it is difficult
to move quickly - trunks of bamboo form almost solid walls, and between them
the large animal simply cannot walk through. But smaller species of rodents,
adapted to existence in this original biotope, prosper. At first it is difficult
to find out about presence of these small mammals in dense bamboo wood, but
gradually these animals find themselves out. They run on wood litter, rearing
from time to time among trunks, and showing improbable, practically squirrel-like
speed, climb up on smooth and firm stalks of this graminoid where even for
cat it would be difficult to seize against something. It is the Hawaiian bamboo
rat, the descendant of black rat (Rattus rattus).
Hawaiian bamboo rats are small rodents, like only the young grey rat by size.
They have kept the appearance typical for representatives of family, but have
got some features, permitting to survive in specific biotope. Representatives
of this genus of rodents meet exclusively in bamboo thickets, and are not found
anywhere more. On each island of archipelago there is the species of these
rats distinguished from others in size and features of coloring.
The Hawaiian bamboo rat dexterously swarms up stalks of bamboo. It has tenacious
claws, but it is not enough to it to show such quickness like it swarms up
trunks of bamboo. On palms and feet of rodent the skin forms small holes acting
like suckers. With their help rat can quickly run on smooth trunks of bamboo.
To improve action of suckers, sudoriferous glands on their edges are hypertrophied,
and allocate stickier secret providing the best sticking to trunk.
The short wool of the rat is colored yellowish – brown. On back of rodent there
are dark longitudinal stripes – they mask this animal in the world of vertical
trunks of bamboo. The amount of strips and their pattern considerably vary
at different individuals.
Hawaiian bamboo rat completely depends mainly on large species of bamboo with
thick trunks. Such choice of habitat is connected with the fact that all species
of Hawaiian bamboo rats arrange shelters, simply gnawing out apertures in ripen
hollow trunks of bamboo. After rats birds use their tree-trunk hollow: on Hawaii
there are no woodpeckers, and, except for rodents, there is nobody able to
make tree-trunk hollows here. And the natural tree-trunk hollows formed in
trunks of some trees, does not suffice for all. Therefore some birds depend
on well-being of bamboo rats.
This rodent has one more important “duty”: because these rats eat young sprouts
of bamboo, they thin out thickets of this graminoid in such way. To gnaw hollows
and to chew firm stalks of bamboo, at this rodent large incisors and short
strong jaws were developed. Cheekbones of this rodent are wide, and jaw muscles
are very strong.
All bamboo rats are solitary animals preserving the territory from neighbours.
Only during breeding season males can come in territory of females for pairing.
Usually per one year at the female it happens three litters of 5 - 6 cubs.
They are born naked and blind, but develop rather quickly. Bi-monthly animals
leave mother, and search to themselves for suitable place for life. Usually
in territory of each adult animal it happens some shelters gnawed in trunks
of bamboo, and young animals frequently lodge in them, choosing ones in which
the lawful owner of territory was not for a long time.
Trunks of bamboo are very strong, and animals settling in them are perfectly
protected from many enemies. But occasionally the population of Hawaiian bamboo
rats has not best times. During flowering of bamboo (one - two times per one
century) number of Hawaiian bamboo rats is considerably reduced: the bamboo
dies off after flowering simultaneously in big territories, and rats sometimes
appear without food among a congestion of died out trunks of these graminoids.
The survived rats in small amount pass to other forage, eating young sprouts
of bamboo, slowing down naturally its distribution. Also at lack of the basic
forage they eat fleshy rhizomes and the bottom part of stalks of other grasses,
including other species of bamboo.
On other islands of the Hawaiian archipelago close species of bamboo rats live:
Black-backed bamboo rat (Bambusiphagomys melanodorsus) lives on Molokai Island.
It is the small species representing by the size something mean between rat
and mouse. This rodent lives on medium-sized species of bamboo and at tops
of large species, choosing the internode of suitable size. The basic colouring
of its wool is sandy – yellow, and on back from a nape up to the basis of tail
one wide black strip stretches. Occasionally on sides there are pale longitudinal
strokes.
Narrow-striped bamboo rat (B. angustizonus) is also small species meeting on
Maui Island in thickets of small-leafed species of bamboo. It arranges shelter
in the basis of trunks in the thickest internodes, but feeds mainly at tops
of bamboo, eating insects and young leaves of this grass. On its back set of
narrow longitudinal stripes of brown color stretches on grayish - white background;
stomach is white.
Half-striped bamboo rat (B. hemifasciatus) lives on Lanai Island. This rat
differs from close species in remarkable colouring: on forward part of body
strips are absent, they begin only on middle of back, and cover waist and hips
of animal. The basic colouring of this animal is light grey with black strips.
On head and cheeks of this rat there are white spots using for recognition
of congeners. Except for colouring, this species differs in more graceful addition:
lengthened body, short forepaws, narrow muzzle and long tail. This rat reaches
usual length of grey rat, but weighs approximately twice less than it.
Hawaiian gatherer rat (Pacifirattus collector)
Order: Rodents (Rodentia)
Family: Murines (Muridae)
Habitat: Hawaiian Islands, rain tropical woods.
Hawaiian islands differ from set of other islands in rather big area and huge
variety of possible habitats given to live creatures. The complex mountain
relief creates a plenty of rather isolated habitats and as the consequence
of it, set of various though also closely related species of small live creatures
lives on Hawaii. They differ from each other in features of appearance, way
of life and behavior. Such variety gets sometimes the extremely fantastical
forms.
Mammal of Hawaiian Islands evolved mainly from animals introduced by people.
And rodents reached the greatest variety among them. Having high speed of alternation
of generations and small size, they could occupy various ecological niches.
Island descendants of rats differ from each other in size, way of life and
features of behavior.
In human epoch in territory of Australia and New Guinea bower birds, and in
pampas of South America large rodents viscachas lived. These animals had differed
in one interesting feature of behavior: having predilection for various bright
objects, they collected them, arranging in special places original “exhibitions”.
The similar creature lives on Hawaii in Neocene epoch. It arranges round wicker
drays in branches of bushes, and places “collections” of shining exoskeletons
of beetles and snail shells around of them on broken off knots. Owners of building,
the breeding couple, from time to time flash in foliage, hastening to the dray
or back.
This four-pawed “collector” is Hawaiian gatherer rat, a creature like large
grey rat (Rattus norvegicus) by size, and by the way it is its strict descendant.
This rodent leads semi-climbing habit of life, swarms up bushes well, and arranges
drays at small height from the ground. For protection against enemies this
rat builds drays in prickly bushes.
Being a day time animal, Hawaiian gatherer rat has the bright colouring appreciably
distinguished from monotonous and dim colouring of ancestor. At this rat there
are red back with longitudinal lines of dark speckles, brown head and white
spots near black ears. Also there are white points above eyes. At this rat
there are large eyes and wide rounded ears through which there leaves surplus
of heat in hot day. Paws are armed with sharp tenacious claws; joints are very
mobile, hand and foot can turn aside to the significant corner.
This rat is omnivorous, but prefers food of animal origin. It eats various
snails including predatory ones, at an opportunity ravages nests of birds and
eats insects. Frequently this rat eats large beetles, and drags their shells
to the dray to supplement its “collection”. This rodent lives at edges and
damaged sites of wood where bushes plentifully grow. It is able to swarm up
branches perfectly, and here prehensile tail making more of half of the general
length of rodent helps the rodent. The tail is naked, covered with grey skin
with separate bristles. It is used as the balance weight more often. Jumping
from branch to another, this rat can drive by tail, or rotates it to turn over
in air and to land on paws like the cat. On the bottom side of tail the cross
horn combs permitting to catch for bark stronger when rat moves in bush, are
advanced.
Dray of this rodent is similar to squirrel’s one: it is spherical, wattled
of rods, arranged in densest part of bush thickets. Similar drays martenrats from Tonga islands plait independently of them, but at the Hawaiian rats, as
against to Tonga ones, the entrance to dray in turned not from below, but sideways.
This rat frequently alters for habitation bird's nests, preliminary having
attacked on owners and having ravaged their clutch. Drays made among prickly
bushes are especially appreciated. This species lodges in drays in pairs kept
for some seasons, and frequently till all life. For strengthening of relations
between partners rats use various symbolical “gifts”, and by their amount near
the dray it is possible to judge force of matrimonial connections: the strong
pair in the prime of life collects many various “gifts”, and successfully protects
them from encroachments on the part of congeners. Usually these rodents collect
bright snail shells under “butcheries” of local hookbill bird eating snails,
and decorate with them an entrance to the shelter. For the organization of
“exhibition” pair of rats gnaws off a part of branches near to bush in which
the dray is arranged, clears them of foliage and bark, and picks on them shells,
dead beetles of bright coloring and if their number is too little – live flowers.
The single male, having won itself a bush suitable for habitation, first of
all organizes “exhibition”, and arranges dray only later. The more bright and
large will be such collection of bright objects, the more obviously force and
dexterity of animal had collected and had kept it. Different pairs have different
predilections - near some drays there is more live flowers, near others shells
or elytrums of beetles are presented. Pairs at gatherer rats are to no small
degree formed on generality of “aesthetic views” of partners. The “exhibition”
is regularly updated: the withered flowers and dried berries are replaced by
fresh ones, and new “exhibits” found, or stolen at neighbours from time to
time are added there.
During courtship displays the male drags in teeth bright objects from “collection”,
showing them to female. It serves as original “substitute” of secondary sexual
attributes. Partners from amicable pair in common steal ornaments at neighbours
and protect “collections” from ravaging. If necessary these rodents can drive
off from bushes even local huge herbivore birds (giant forest geese), frightening
them by shrill squeal. If birds do not recede, rodents even attack them, biting
birds in unprotected paws. Some pairs of these rodents can operate in common
against the large enemy.
Seasonal prevalence in breeding of the Hawaiian gatherer rats is not present.
At any time year in drays of these rodents it is possible to find cubs of various
ages. In litter of this rodent three - four times per one year up to eight
- eleven naked blind cubs appear. The female cares after them, leaving posterity
only for feeding, and the male is completely occupied with protection of fodder
territory and “collection” from neighbours. At week age the young growth becomes
covered by wool, for the twelfth day starts to see and hear. Up to monthly
age rat cubs stay in dray, and later start to play already outside of home:
swarm up branches, drag small objects, trying to steal them at each other.
Bi-monthly cubs already receive punishment from parents, and one by one abandon
parental territory. In youth, not having an own home, they are very vulnerable,
and at this time perish from various predators. Up to half-year age, when the
young animal becomes strong enough to win to itself territory, only few individuals
survive. Life expectancy of these rodents is short – only few animals live
till three years, and the majority does not live even till two years.
Mangrove builder rat (Architectorattus piscatorius)
Order: Rodents (Rodentia)
Family: Murines (Muridae)
Habitat: Meganesia, mangrove forests of Arafura Lake.
Rodents were only terrestrial placental mammals in Australia before the human
arrival to this continent. Under human influence the fauna of Australia had
substantially lost the originality, and a plenty of non-native species introduced
from other continents had appeared in it. During the time pased from the moment
of extinction of mankind, some lines of placentals had declined, and others,
on the contrary, continued the prospering. Rodents are especially succeeding
group among them. The type of anatomy, characteristic for rats and mice, appeared
very successful, and numerous rats had settled in various biotops of Meganesia.
One of their species due to versatility of behaviour had developed mangrove
forests separating by thousand-kilometre wide barrier Arafura Lake from the
ocean. It is very specific environment where small islets of land alternate
with impassable marshes and rich mattings of roots and branches of mangrove
trees.
The large species of rats well adapted to life in such inhabitancy lives in
branches of mangrove trees. Its settlements represent “towns” of several tens
of nests looking like small huts arranged on branches. Because of this feature
the animal is named mangrove builder rat.
This mammal is medium-sized aquatic rodent: its body length is about 30 cm.
It has anatomy typical for rats, long flattened muzzle and short forepaws with
mobile fingers and sharp claws. Rear legs are big and strong, and have membranes
between toes. Auricles are small. When rat dives, acoustic duct is closed by
the special skin valve. Mangrove builder rat has long tail, which serves as
a counterbalance when it swarms up branches. Wool of animal is velvety and
rich, but short. This rat greases it with fat secretions of special glands,
and spends a lot of time cleaning it. Cleaning of wool at these animals is
a mark of mutual attachment, and the individuals standing below at the scale
of rank express submission to stronger individual this way. The top part of
body of mangrove biulder rat is colored beige; throat and breast are white.
This rat eats fish, crabs and insects, and also consumes fruits and young leaves
of mangrove trees. It is able to swim and dive well, and during the inflow
it hunts for fish and other water animals. Swimming under water this rodent
presses forepaws to breast and rows by hinder legs. Mangrove builder rat leads
social habit of life and settles among mangrove thickets in colonies numbering
some tens of individuals. The inhabiting in holes and hollows traditional for
rats does not fit to conditions of mangrove forest – ground is regularly flooded
here, and hollows in tree trunks are not so extensive to contain the numerous
colonies of animals. Therefore during the evolution at these rats the special
form of building behaviour had developed. Each breeding pair constructs for
itself the original dwelling – primitive conic-shaped “tent” which is located
above a level of the highest inflow. By this feature mangrove builder rat is
similar to martenrat from Tonga Islands – the rodent developed during the evolution
process the similar kind of shelters independently of it. In the basis of “tent”
there is a small wum platform made by animals of fibrous leaves of palm tree
and pandanus, or of bark of mangrove tree sprouts. Above it the dome of arched
bent rods is erected, on which the roof made of leaves is attached. The ready
construction has conic shape. Houses of mangrove builder rats form a kind of
“village” where up to ten pairs of adult animals and numerous young ones live.
At the exhaustion of fodder resources the colony abandons “village” and moves
to new places where new constructions are quickly made.
Within one year the pair of adult rats may bring into the world up to four
litters of 6 – 8 cubs. They develop quickly and already at the age of 6 – 7
weeks become independent and make their own constructions at the edge of “village”.
At the age of half-year they become sexually mature. Life expectancy at these
animals does not exceed three years.
Hawaiian mole mouse (Xenospalax subterraneus)
Order: Rodents (Rodentia)
Family: Pacific mole mice (Xenospalacidae)
Habitat: Hawaii (Big Island), ground in valleys and woods.
Before human occurrence at Hawaii mammals were absent at islands and only insects
and their larvae could occupy an ecological niche of underground digging animals.
But historical colonization epoch the fauna of islands had replenished with
various animals, alien to indigenious fauna on fault of people. At Hawaii various
species of ground mammal, which began to develop ecological niches inaccessible
to birds, had appeared. Moles and other digging animals were not introduced
to Hawaii, but their place uniquitous rodents had occupied: they are descendants
of mice avoiding a competition to descendants of two species of rats, also
introduced to islands by people.
The ground in woods of Hawaiian Islands is penetrated with set of underground
burrows in which their builders - completely blind rat-sized animals – run.
By body shape they are similar to mole rats (Spalax) living in Eurasia, but
they are descendants of house mouse. Therefore they are named Hawaiian mole
mice. They differ from true mole rats in features of behavior: if true mole
rats frequently left on the ground surface for feeding, their Hawaiian “doubler”
does it extremely seldom, eating only roots and tubers of plants.
Hawaiian mole mice are similar to continental species by features of anatomy
developed at both groups independently. At them there are the same cylindrical
body, short paws and large head. Main digging adaptations of Hawaiian mole
mouse are its head and strong incisors. Skull of this animal is wide and short,
and incisors of top jaw are very big. Lips of animal densely close mouth from
the ground by special outgrowths, not impeding incisors. Occipital muscles
are very strong, for their attachment on back edge of skull of the animal the
special crest stretches. Muscles lowering head are also strongly advanced.
During the ground digging this small mammal lifts head making effort and ramming
a vault of formed hole, and then lowers it, ploughing by incisors deep furrow
in the ground before itself. Animal rakes away the loosened ground by forepaws
under stomach and pushes it out from hole, or simply stamps by paws.
This animal is completely blind, and its eyes are reduced even in greater degree,
than at moles – eyeballs are abscent and the optic nerve had reduced. The top
of head and shoulders of Hawaiian mole mouse are covered with thick cornificate
skin – animal forms a vault of hole by them. Corneous armour on head closes
the place where ancestors of this animal had eyes, and also covers nose from
above. It is additionally strengthened with bones of skull of animal grown
together in continuous osseous vault.
Because paws do not accept direct participation in digging, they are rather
short. Animal only rakes away the friable ground back by them. Tail of Hawaiian
mole mouse is also very short and hairless. Animal is covered with velvety
fur of black color. On muzzle of Hawaiian mole mice long whiskers, with which
help animals are guided in space of underground courses, grow.
Hawaiian mole mice build extended systems of burrows in wood, and arrange the
special drainage tunnels through which the water leaking to burrows because
of daily rains is removed from them. As animals are vegetarians, the competition
between them is not as sharp, as, for example, at moles. Hawaiian mole mice
eat roots and tubers of plants, and even parts of such species which are not
eaten by other animals.
Hawaiian mole mice breed practically the year round. For one year the female
can draw up to five hatches numbering 6 – 8 cubs. They grow quickly, and at
monthly age already begin independent life. At three-monthly age they already
can give birth to posterity. But, despite of such high rate of breeding, the
significant amount of young growth perishes: when the colony appears overpopulated,
the part of its inhabitants (mainly young animals) feels stress. It occurs
because of too big frequency of meetings between animals, and less strong animals
aspire to leave earlier safe holes. At this time animals frequently settle
not in systems of holes, but on the ground surface. It happens that such animals
movable by aspiration to avoid of stress appear on ground surface in day time,
run in wood, not trying to hide, and at times even try to force rivers. The
significant part of these of “victims of stress” perishes, but some ones succeed
to leave far enough and to found a new settlement.
Under the ground one of main dangers – the descendant of intruder like them,
the large digging blind snake catches Hawaiian mole mice. And on the ground
surface local predatory birds, and even descendants of rats catch them.
At the next islands of the Hawaiian archipelago there are close species of
these rodents:
Mole mouse of Lanai Island (Xenospalax lanaiensis) is the small species (equal
to large mouse in size), distinguished by spotty colouring: on light grey background
there are dark spots. Separate individuals of this species are almost completely
black. This species has similar biology, but spends more time on ground surface.
At night it leaves to feed among bushes.
Mole mouse of Maui Island, or half-naked
mole mouse (Xenospalax mauiensis) differs
from other species in strange appearance: the fur of yellowish-grey color had
remained at this species only on the bottom part of body and on shoulders.
Head, back and sides of this rodent are hairless, covered with rough grayish
skin. This rodent lives in warm rainforests, and is able to swim well. The
young growth can even settle during floodings.
Giant
mole mouse (Xenospalax giganteus) lives at small Kahulawi Island near southwest
coast of Maui Island. It is the largest species of genus: the length of body
reaches 25 cm. It differs in greater, than at other species, propensity to
leave on ground surface. Its wool is colored rusty-brown color, and corneous
“helmet” on head is rather small.
Giant
wood porcupine (Aepythizon megatherioides)
Order: Rodents (Rodentia)
Family: Wood porcupines (Erethizontidae)
Habitat: woods at plains and mountains of northwest of Northern America (to the
west up to southern areas of Beringia)
After extinction of plenty of hoofed mammals species at the border of Holocene
and Neocene other herbivore mammals had occupied become empty ecological niches
having caused fantastical species. There were rodents among them, huge by variety
order including in Holocene basically small and absolutely tiny species. Certainly
in various epochs of Cenozoic rodents more than once “tried” to come to middle-
and large-sized class, and sometimes it happened most advantageously: the rhinoceros-sized
rodent lived in South America and Pleistocene North-American species of beavers
reached the size of bear had appeared among them. In Holocene largest representative
of this group were beaver and capybara lived in rivers. But in Neocene having
occupied various ecological niches rodents have fairly increased their size.
In South America among rodents forms similar to antelopes and pigs, and also
giants similar to ground sloths of Pliocene and Pleistocene have appeared. At
the territory of Mexican plateau the bear-sized
porcupine has lodged in Neocene. But in wood zone of Northern America this
porcupine has even larger relative.
Giant wood porcupine is very large representative of Neocaenic rodents (it weights
up to 1 ton; growth of animal standing on hinder legs is up to 3 meters, general
length is over 5 meters including tail). Back legs of this animal are longer
than front ones; due to it animal can stand on two legs eating plants. Feet of
hinder legs are covered with thick cornificate skin therefore animal equally
easily can walk both on stones heated up by sun and on winter ice. It is a herbivorous
animal; its food includes leaves and branches of various deciduous trees. In
winter giant wood porcupine can eat even poorly nutritious branches of coniferous
trees and roots digging out from ground. On forepaws of porcupine there are curve
claws with which help animal gets food to itself and effectively protects from
enemies. When the animal moves on four legs, it leans against lateral surface
of fingers, claws thus are directed by tops to each other.
This species of animals lives in temperate zone frequently rising highly in mountains.
Therefore in winter it should endure rather strong frosts. Fur helps it to do
it becoming to winter thicker and denser. Colouring of fur is changeable, as
well as at its desert neighbour: there are variants of colouring from light brown
up to grey and straw-coloured. In fur the main protective weapon of the huge
wood porcupine is hidden – very thick (up to 2 cm at the basis) needles up to
40 - 60 cm long. They are especially numerous on neck (from above and from sides)
and shoulders and then grow by strip along back (up to the basis of tail). Thus,
all places which predator prefers to attack are reliably protected. Needles are
mobile; animal can lift and lower them depending on a situation. Sometimes (especially
if the predator is weak or inexperienced) giant porcupine itself can attack it,
striking impacts by needles growing on shoulders. Besides it can strongly bite
by huge incisors. During fight the porcupine roars for frightening. But usually
adult animals seldomly show their abilities: while they are strong and healthy
they practically have no enemies. And cubs of huge porcupines can become easy
catch of predators departing far from mother.
At the female in pack there are 1 – 2 cubs. The new litter appears once a year,
but young growth long time stays with the female: last year's cubs continue to
feed with mother after birth of new ones, abandoning her up to the time of sexual
maturity (at the third year of life). Cubs are born without needles being overgrown
by them at the first month of life. But the newborn wood porcupine is covered
with wool, can see and already in half an hour after birth studies to walk. The
wool of cubs of the first year of life is dark but after the first winter, during
spring moult, colouring is shown which the animal will have all next life.
Sexual maturity comes at the third year of life. At this time young animals are
already independent, and take part in courtship games which take place at the
end of summer. On borders of individual possessions battles of males for the
right of pairing with females happen. But these tournaments are not as substantial
as at the desert bear porcupine: usually two males battle for one or two females.
But fights happen fierce seldomly: more often all events may end by might demonstration.
Contenders peel bark on tree trunks, pull out bunches of grass from the ground,
urinate trees plentifully and snort loudly. Females observe of the duel from
the side stimulating contenders by characteristic rumbling and spraying strongly
smelling urine.
Pairing proceeds not for long and animals miss soon. But during carings the excited
males can take cubs of any female for contenders and attack them. Frequently
grown up young growth survived in winter and has got stronger, perishes from
attacks of males in courtship season. But porcupine has successfully survived
in youth can live about 60 years.
Barocavia
(Barocavia potamophyla)
Order: Rodents (Rodentia)
Family: Capybaras (Hydrochoeridae)

Habitat: rivers and bushes of South America.
During the most part of Cenozoic South America was isolated continent. Its
flora and fauna developed independently from other continents and has reached
high level of endemism and originality. In late Cenozoic connection of South
and North America was restored and as a result of immigration of North-American
species the most part of South-American ones appeared in conditions of isolation
and not taking place similar “durability tests” has died out. But some groups
have remained rather widely spread and various – for example, characteristic
South-American rodents. When in Neocene South America appeared isolated from
world again around by Panama passage, these animals became basic forms of local
herbivores having evolved numerous and various species having ecological analogues
in other parts of the world: original “doubles” of antelopes and pigs. And
the place of hippopotamus in rivers and lakes of South America was occupied
by giant rodent – huge barocavia, the descendant of capybara.
Barocavias are widely settled in rivers of South America. In constant reservoirs
they live settled life, and in seasonal drought areas had turned to nomades
and make migrations to deeper rivers and lakes. Usually these huge animals
spend almost all day in the river eating or having a rest, and in the evening
come to bank meadows to graze.
The barocavia is huge semi-aquatic rodent resembling hippopotamus externally
and by habit of life. It grows to 3 – 4 meter length (male is larger) at height
at a shoulder up to one and half - two meters. The adult male of this species
can weigh up to three tons. At barocavia there is thick skin covered with thin
rough wool of brownish color. Tail at this giant rodent is not present.
Head of barocavia is disproportionally big, having big cheek-bones, and heavy.
On the top jaw of animal long rigid whiskers grow. Incisors in bottom jaw are
wide and shovel-like. They are directed forward and serve for raking of floating
plants from water surface and for excavation of roots from bottom or coast.
Top incisors are covered with white enamel, their length reaches 20 cm. They
permit to have a cut through even trunk of young tree easily, and in case of
predator attack help to give them adequate repulse. Molars of barocavia are
wide and knobby. It is connected with diet of animal: barocavia consumes for
day plenty of enough soft and sappy vegetative food and chews the most part
of day.
Small eyes and short rounded ears of barocavia are shifted to the top part
of head as at hippopotamus. At this animal there is bad sight but keen sense
of smell and excellent hearing.
Legs of barocavia are plantigrade, thick and short. Toes are united by common
cover of soft tissues and form the structure similar by form to the elephant
foot. Under them the pillow of elastic tissue impregnated with fat allowing
an animal to walk easily, springly and silently despite of apparent awkwardness
is located. Claws (on forward leg their number is four, on back one - three)
are thick and hoof-like. Barocavia can swim and dive well rowing by legs. Under
water animal walks easily and even is to some extent graceful, slightly touching
a bottom. In case of necessity these animals easily can cross rivers some kilometers
wide.
Barocavias live in big herds numbering up to 30 – 40 individuals. Such large
herd is non-uniform: in it there are some breeding groups – one adult male
for 3 – 5 females and their cubs of first year of life. Herds of smaller size
are, as a rule, groups of bachelor males not having families by different reasons.
All breeding groups in herd are approximately equal by rank. In case of danger
herd can unite and defend from predator collectively. Barocavias try to drive
off ground predators widely opening mouth and showing huge teeth and light
pink gums. Thus adult animals abruptly bellow and growl. Water predators - large
carnivorous fishes – as a rule, are not dangerous to barocavias, though
they can drown and eat newborn cubs. At the case of fish attack adults protect
cubs by bodies, rear and fall to water, splashing fountains of water and trying
to trample attacking fish. Sometimes adult animals try to bite fish or to snap
it and to cast ashore (according observations of zoologists African hippopotamuses
of Holocene epoch acted this way).
In rainseason when rivers are widely overflowed and transform significant areas
to marshy plains, barocavias leave main channel of river and go to near lakes
and bogs. Here they feed with marsh plants plentifully expanding on damp ground,
supplementing the diet by branches of trees and bushes. Each herd occupies
the certain territory which edges are marked by heaps of manure. By smell of
manure any neighbour can know as great the herd is, what animals make of it,
and as frequently it visits this place.
These rodents spend hottest time of day in water having a rest and eating water
vegetation. During feeding barocavia sticks teeth under reed turf, pulls out
the whole sheaf of stalks by strong movement of head upwards, “rinses” it in
water by lateral movements of head and then starts to chew phlegmatically.
These rodents to sleep prefer on land in places where ground predators can
not reach: frequently “bedrooms” of barocavias are situated on small islands
among bog or on sandy islands in channel of river. Dream time at the barocavia
lasts from second half of night till late morning.
The role of barocavia in ecosystems of South American rivers is great: eating
plants they interfere river overgrowing and bogging, and on their manure phytoplankton
– basis of efficiency of river ecosystems – plentifully reproduces. Eating
bushes on river banks barocavias promote development of graminoid communities
serving for feed to local running rodents deermaras and giant pacas.
The breeding season at barocavias begins approximately in middle of rainseason:
thus cubs will be born in the most favorable season, and the probability of
their survival will be much higher. Out of breeding season males practically
do not notice each other and even can graze grass side by side. But when the
female from breeding group shows the readiness for pairing, the male becomes
terrible and jealous husband. Bassy uterinely roaring it snatches to any one
in which competitor will be seen. During intraspecific duels barocavias do
not use huge teeth and simply push each other by shoulders and strike impacts
by forehead or lateral part of head.
Caring for the female male slightly pushes her head, puts his head to her back
or neck. Thus he “coos” – utters special abrupt humming sounds, and walks “on
tiptoe” highly lifted head up and trying to seem even larger. If the female
is not ready to pairing, she keeps head to the caring male and from time to
time quietly growls showing incisors. Accepting male carings female nestles
against his side, “coos” in unison, and sniffs head of male from below. Pairing
at massive barocavias occurs in water.
Pregnancy lasts about eight months, and once a year female gives rise to two
well-advanced cubs with opened eyes, covered with wool. The newborn barocavia
weighs about 50 kgs. Already at the first day of life the cub can walk and
even tries to swim. At fortnight age young barocavia tries to eat grass and
completely passes to feeding by plants at the age of three months. At this
time young animal can weigh already up to 300 kgs. Barocavia becomes completely
adult at the age of about two years, and life expectancy can be about 30 -
40 years.
Rafter
coypu (Paracoypu totorae-lacustris)
Order: Rodents (Rodentia)
Family: Coypus (Myocastoridae)
Habitat: Totora Lake, Andes.
Along Pacific coast of South America the extended mountain circuit – Andes
– stretches. Conditions of life in this area sharply contrast with an environment
of rainforests of Amazonia or savannas of Gran Chaco. The high mountain world
is an empire of cold, sharply expressed daily and seasonal change of temperatures
and superfluous sunlight. In ice age the high-mountainous climate was dry and
very cold. In Neocene the climate began more humid; the increase of amount
of rains had caused occurrence of large lake at Altiplano plateau. In human
epoch at this plateau there were some lakes among which Titicaca lake was the
most known one. Gradually all lakes of Altiplano had merged together, and at
the plateau the huge Totora Lake appeared.
Coypu reached significant evolutionary success in South America. Among descendants
of this animal there is a unique group of mammals – algocetids, huge aquatic
animals. It is the most deviated and specialized branch of rodents making the
separate order. Other descendants of coypus are less specialized, but it does
not prevent them to prosper in nearwater biotops. In South America they are
analogues of beaver and muskrat, and settle down to the cold south of continent.
One descendants of coypu has developed high mountains, and its population lives
at Totora Lake. It is rafter coypu, animal leading unusual way of life and
differing in advanced building behaviour.
Rafter coypu is large species of aquatic rodents keeping anatomic features
of the ancestor. The velvety fur with small amount of guard hair serves to
this species as fine thermoisolation in very cold water of the lake. In fur
colouring significant polymorphism is observed – among animals there are individuals
from grey and straw-coloured up to brown and black ones. Rafter coypu carefully
looks after fur, and combs it by claws for a long time. Forepaws at this animal
have mobile fingers and are adapted to manipulation with small objects.
Ears of this mammal are short, and eyes are small. This animal receives a significant
part of information about the world around with the help of smell sense. On
a muzzle of rafter coypu numerous whiskers grow, helping to orientate in darkness.
Main moving organ of this animal at the swimming is its tail. It is wide, rounded
and looks like cut off at the back edge. Tail is covered with hairless tight
skin and contains the advanced fat layer; therefore warm does not leave an
organism of animal through naked skin of tail. In water rafter coypu swims
with the help of tail, having pressed paws to body and making wavy bends by
backbone. During feeding at shoaliness animal swims slowly, with the help of
webby hind legs. Rafter coypu is able to dive to the depth up to 10 meters
and stays under water till about 5 minutes. But it prefers to use this skill
only in case of extreme danger, because it feeds in coastal shallow zone.
This species is strictly herbivorous animal, eating rhizomes of totora reed
(in language of Indians of Andes it is the name of the reed known in human
epoch, hence the name of lake) and tubers of water plants. Eating rafter coypu
simultaneously receives building material for its shelters. This animal digs
out sprouts of reed with the help of strong incisors and claws of forepaws.
At the dug out plants rafter coypu bites off and eats rhizome and the bottom
part of stalks, and pulls a bunch of poorly nutritious stalks to original shelter
in which it spends night.
These animals build rafts and huts on them of reed. To construct a raft, rafter
coypu bites off stalks and branches of coastal plants, drags them to common
heap floating on surface of water, and constantly renews the construction.
As a rule, some animals begin simultaneously construction of raft. They constantly
drag the cut off rods and stalks of reed on rafts, and trample them down, moving
on rafts. On such raft the hut is placed, in which the pair of adult animals
and their posterity lives. Hut has the hemispherical shape, reaches one meter
height and has thick friable walls. The entrance opens sideways; if the raft
is great enough, near to an entrance animals specially make an outlet to the
water and clear it regularly, dragging off from it the rests of plants.
Small rafts, constructed by separate family groups, may gather to form big
floating islands and give a haven for other creatures. When some rafts are
linked together, their inhabitants do not try to uncouple them – rafter coypus
are sociable enough and between them there are no serious conflicts. They simply
throw more reed stalks on edges of linked rafts and form unite large raft.
Running on it, animals trample down its surface, and the rests of their food
serve as a building material. Such rafts may exist for many years, constantly
being renewed. The part of reed rhizomes not eaten by rafter coypus sprouts
in thickness of rotten stalks, and additionally strengthens the construction.
Sprouts of reed make the way through surface of raft, and serve as good top
dressing for these animals. In the middle of raft animals eat sprouts completely,
and on edges the bitten stalks of totora reed plentifully branch, forming green
fringe.
The pair of adult animals rises within summer one litter – up to five cubs.
The young growth of these animals is born with open eyes and covered with rich
wool. The female cares of posterity within two months, and at this time male
is engaged in repair and construction of raft alone. Young animals leave dwelling
already at week age, and monthly cubs swim perfectly. They live in common with
parents within the first winter. At this time young females reach sexual maturity,
and males become large enough (they mature later). Females leave parental raft
and settle on rafts of other males, forming pairs with them. Young males are
banished by their own father, and they start to build their own rafts or occupy
and renew empty huts.
The idea about existence of this species was proposed by Momus, the forum member.
Chilean algocetus (Algocetus chiliensis)
Order Algocetiforms, “algae whales” (Algocetiformes)
Family Algocetids (Algocetidae)

Habitat: Pacific coast of South America.
Human activity and climatic changes at the boundary of Holocene and Neocene
had caused fast disappearance of plenty of species of sea animals. Cetaceans,
pinnipeds and sirens had become extinct irrevocably till this epoch. But after
mankind disappearance and stabilization of climatic conditions life began to
restore the lost positions and to form new ecosystems again. Seas of northern
hemisphere in Neocene are occupied by birds – gannetwhales and flightless loons.
These animals dominate in Atlantic and Arctic oceans. In Pacific Ocean large
flightless auks hatchetbills replace them, and at the Asian coast algal geese
and penguin duck also live. But in cold and temperate waters of southern hemisphere
birds are presented only by flightless penguigulls of several species. These
birds eat animals, and mammals came to the place of herbivorous marine animals.
Sea mammals of southern hemisphere descend from South America. They are descendants
of coypu (Myocastor coypus) widely settled in a moderate and subtropical zone
of continent. Coypu had successfully passed the period of anthropogenous pressure,
and even had widely settled due to human activity – it was domesticated.
Sea mammals of Neocene are herbivores. In early Neocene when sirens – completely
aquatic herbivorous mammal – had disappeared, evolution of coypu had gone especially
roughly. Till some millions years descendants of coypu had passed approximately
the same way, than cetaceans in early Eocene epoch did – from small semiaquatic
creatures up to completely aquatic large mammals lost connection with land.
Descendants of coypu form the separate order of mammals – Algocetiformes, “algae
whales”. In their anatomy features of rodents and features of obligatory aquatic
mammals (whales and sirens) developed convergently are combined. Algocetids
are large animals; smallest of them reach the length of 3 meters, and at coast
of Antarctica and at the south of South America the giant about 7 meters long,
weighting over 6 tons, lives. They resemble sirens externally a little; it
is the result of similar habit of life.
The muzzle of algocetes differs in advanced, mobile and partly doubled upper
lip on which bottom side lines of corneous combs grow. With the help of such
lip the animal easily scrapes off from stones seaweed which it eats. The bottom
jaw differs in massiveness, and incisors in it are very large and wide. Such
incisors help to tear off from stones not only seaweed, but also sedentary
animals – mollusks and barnacles.
Nostrils of algocetids are shifted to the top part of head – such structure
permits the animal to breathe, having put out from water only small part of
head. Large individuals emerge for some minutes, and then can hold the breath
till half an hour. Cubs emerge to breathe much more often.
Though algocetids are completely marine animals, not capable to leave on land,
they swim not so well. These animals prefer to move in shallow water similarly
to Steller’s sea cow – making steps on bottom by well advanced mobile forward
extremities. On them two large claws (on II and III fingers), assisting to
cling to bottom, are present. IV and V fingers are reduced, lack of mobility,
and only support a plane of flipper. The thick layer of fat serves not only
thermoisolation, but also a fine float, reducing loading to extremities of
animal. Rear legs of algocetids had reduced completely, and on the tip of tail
the wide rounded fin is developed. Algocetes are sluggish swimmers, and during
storm they prefer to hide in rich thickets of seaweed which soften force of
waves.
Eyes at algocetes are small, but sight is rather quite good. Animals prefer
to live in places with transparent water, supporting visual contact to each
other. They also can produce sounds with the help of air bladders in nasal
ducts. External auditory meatus at algocetids is not closed, but an eardrum
is thick. Bones of internal ear are isolated from skull by special fatty tissue,
like at cetaceans. The upper lip is covered with thick whiskers (hair thickness
is up to 3 mm), which help to search for food in muddy water.
Algocetes keep in families including a breeding pair or the male and two females,
and their posterity of last two – three generations. At the largest species
living near Falkland Islands and in Antarctic Region, herds of several tens
of individuals including some breeding groups are formed. Pregnancy lasts about
16 months at large species and about 11 months at smaller ones. As a rule,
the female at all algocetid species gives rise to two well advanced cubs. Only
at the largest species there is only one cub. Cubs are born covered with thin
fur which drops out till first days of life. At the delivery the female is
surrounded by neighbours push the cub out to the surface of water. When the
cub starts to move, they leave it, and only its own mother cares of it. But
if necessary, females of algocetuses protect young growth against large predatory
fishes together. Nipples of the female are located on sides of body, approximately
at the level of pelvic girdle hided in thickness of muscles. To make more convenient
to cub to be fed, the female lies down on the side, and the cub sucks milk,
having put nostrils out to the surface of water, and breathing. Cubs suckle
milk about half-year, but already from bi-monthly age start to eat food of
adult animals.
In Neocenic fauna algocetids are presented by one family and four species.
Chilean algocetus lives at Pacific coast of South America, and the separate
population with doubtful specific status inhabits waters of Galapagos Islands.
The length of this species reaches 4 – 4,5 meters. This species lives in coastal
zone, preferring areas of coast protected from storm waves by barrier of small
islands. Its distribution is closely connected to cold current flowing from
coast of Antarctica. These animals eat brown and green macroalgae. Due to their
fast growth animals lead almost settled way of life, moving within the borders
of the territory to some kilometers. This species willingly swims to lower
reaches of short rapid rivers flowing down from Andes to Pacific Ocean. In
such way these animals escape of ectoparasites perishing in fresh water.
Patagonian coypocetus (Coypocetus patagonicus) inhabits rivers of Atlantic
coast of South America. It is the smallest species of algocetes – the length
of adult individuals hardly exceeds 3 meters. It constantly lives in fresh
water and eats aquatic flowering plants. At this species there are rather large
claws – with their help the animal digs out rhizomes of water plants and protects
itself from enemies. Its forepaws are mobile; with their help animal can bring
food to mouth. Digging out silt and eating water plants, these animals interfere
with bogging of the rivers. To wintering these animals migrate from upper streams
and move down to the mouth of the river, forming there big congestions destroying
water vegetation along many kilometers of riverbanks. At the lack of forage
they may swim for the short time to coastal sea waters.
Closely related to the previous species coastal coypocetus (Coypocetus
littoralis) has wide area: it lives along the Atlantic coast of South America. This animal
does not meet in fresh water at all. Its length is about 4 meters. Coastal
coypocetus is rather heat-loving, and does not settle far to the south. But
it also does not love very warm water; therefore the northern border of its
area does not reach mouths of Amazon and Hyppolite rivers.
All species of algocetuses described above live in shallow waters and do not
swim far from coast. Therefore Falkland paralgocetus (Paralgocetus
falklandicus),
the largest species of algocetes, looks the exception among them. It is the
true giant among algocetes, reaching the length of 7 meters. It lives at the
coast of Falkland Islands and rather frequently leaves in open ocean. This
animal frequently feeds in congestions of floating brown seaweed in southern
part of Atlantic. It differs from other species of algocetes in very big fat
incrustation expanding on forward part of head between nostrils and upper lip.
This pillow is used as ram during intraspecific duels. At this species there
are small dulled claws on forward fins, and sometimes the claw on II finger
is completely reduced. A tail fin of Falkland paralgocetus is triangular-shaped,
but it is not such wide, as at whales.
Due to the size and thick layer of fat (up to 15 cm) it is the most cold-resistant
species of algocetids, reached the certain success in evolution. Rather recently
(according the geological measures) the small population of this species migrated
to coast of Antarctica, and later descendants of first settlers settled generally
coasts of continent. The Antarctic representatives of this species became more
omnivorous, and at polar night marine invertebrates frequently make up their
food.